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Contents <br />Page <br />Abstract ................................................. 1 <br />Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 <br />Endangered Fishes of the Upper Colorado River Basin .. ... . .. ......... .. .. . 3 <br />Purpose and Objectives of the Study . .. .. . .. ... .. . ....... .. ... ... 3 <br />Methods and Approach ... . .. .. . .. .. ..... . ... .. . . . . .... .. .. . ... . 5 <br />Review of Information . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 <br />Peer Review and Schedule . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 <br />Ecological Context for Instream Flow Analysis . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 <br />River Ecology and Effects of Regulation on the Endangered Fishes of the Upper <br />Colorado River Basin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 <br />Ecology of the Endangered Fishes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 <br />Dynamic Relationships Between Flow, Channel Geomorphology, and Food Webs . . . . . . . . 11 <br />Influences of Stream Regulation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 <br />Conclusions Based on Review of the Ecological Literature Pertaining to the <br />Endangered Fishes and the Regulation of Flow . . . . . . . . . . . . . . . . . . . . . . . 24 <br />Derivation of Flows Currently Recommended to Protect the Endangered Fishes . . . . . . . . . . 26 <br />Review of Instream Flow Methodology . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 <br />Flow Regimes Recommended to Protect and Enhance Endangered Fishes in the <br />Upper Colorado River Basin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 <br />Problems With the Flow Recommendations of the U.S. Fish and Wildlife Service . . . . . . . . 31 <br />Critical Uncertainties in the Recovery Program With Respect to Provision of River <br />Flows to Protect Endangered Fishes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 <br />Critical Uncertainties at the Information (Scientific) Level of Organization . . . . . . . . . . . 35 <br />Recommendations: An Ecosystem Approach . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 <br />Implement Interim Flows That Reestablish Seasonality . . . . . . . . . . . . . . . . . . . . . 36 <br />Provide Common Understanding of Water Availability . . . . . . . . . . . . . . . . . . . . . . 37 <br />Improve the Standardized Monitoring Program: Add a Community Ecology <br />Perspective . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38 <br />Diversify the Research Program to Resolve Critical Uncertainties . . . . . . . . . . . . . . . . 39 <br />Implement a Peer Review Process . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 41 <br />Implement Adaptive Management . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 41 <br />Cited Literature .. . .. .. . . ... .. . . . . ... .. . . . . . . .. . ... . . .. .. . . .. 42 <br />iii