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Technical Report Series National Biological Survey The National Biological Survey publishes five technical report series. Manuscripts are accepted from Survey employees or contractors, students and faculty associated with cooperative fish and wildlife research units, and other persons whose work is sponsored by the Survey. Manuscripts are received with the understanding that they are unpublished. Manuscripts receive anonymous peer review. The final decision to publish lies with the editor. Editorial Staff MANAGING EDITOR Paul A. Opler ASSISTANT BRANCH LEADER Paul A. Vohs WILDLIFE EDITOR Elizabeth D. Rockwell FISHERIES EDITOR James R. Zuboy TECHNICAL EDITOR Deborah K. Harris VISUAL INFORMATION SPECIALIST Constance M. Lemos EDITORIAL CLERK Donna D. Tait Series Descriptions Biological Report ISSN 0895-1926 Technical papers about applied research of limited scope. Subjects include new information arising from comprehensive studies, surveys and inventories, effects of land use on fish and wildlife, diseases of fish and wildlife, and developments in technology. Proceedings of technical conferences and symposia may be published in this series. Fish and Wildlife Leaflet ISSN 0899-451X Summaries of technical information for readers of non- technical or semitechnical material. Subjects include topics of current interest, results of inventories and surveys, management techniques, and descriptions of imported fish and wildlife and their diseases. Fish and Wildlife Research ISSN 1040-2411 Papers on experimental research, theoretical presentations, and interpretive literature reviews. North American Fauna ISSN 0078-1304 Monographs of long-term or basic research on faunal and floral life histories, distributions, population dynamics, and taxonomy and on community ecology. Resource Publication ISSN 0163-4801 Semitechnical and nonexperimental technical topics including surveys; data, status, and historical reports; handbooks; checklists; manuals; annotated bibliographies; and workshop papers. Copies of this publication may be obtained from the Publications Unit, U.S. Fish and Wildlife Service, 1849 C Street, N.W., Mail Stop 130, Webb Building, Washington, DC 20240 (call 703-358-1711), or may be purchased from the National Technical Information Service (NTIS), 5285 Port Royal Road, Springfield, Virginia 22161 (call toll free 1-800-553-6847).