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INSTREAM FLOWS TO ASSIST THE RECOVERY OF ENDANGERED FISHES 11 <br />Squawfish distribution in Colorado River <br />(1991-1992) <br />50 <br />t 40 - <br />35 <br />3 <br />E 30- <br />Z <br />Z <br />20 <br />10--- <br />8 1 <br />10 <br />4 2 <br />15-mile sections of river <br />Squawfish average length in Colorado River <br />(1991-1992) <br />600 <br />E <br />E <br />500 <br />m <br />J <br />400 <br />300 <br />200 <br />15-mile sections of river <br />Dynamic Relationships Between Flow, <br />Channel Geomorphology, and Food Webs <br />Distribution, abundance, and life histories of the <br />endangered fishes seem to be strongly influenced <br />by availability of physical habitats that are created <br />and maintained by flow dynamics in time and space <br />(Fig. 6). Indeed, squawfish only spawn on clean <br />cobble on specific bars in the sediment-laden river <br />segments of the Upper Colorado system. Hence, a <br />fundamental process-response relationship in- <br />volves the movement of the fish to the bars in <br />concert with flows that first form the bars and then <br />flush sediment off of cobble substratum so that the <br />fish can spawn successfully (Fig. 2; Tyus 1990; <br />Harvey et al. in press). Humpback chub primarily <br />occur in eddies and other hydraulically complex <br />habitats found in constrained channels in the <br />steeper gradient segments within canyons (Fig. 2; <br />700 <br />Fig. 5. Distribution of squawfish by size <br />and number caught in the Colorado <br />River from the Green River confluence <br />(km 0) to the Grand Valley diversion <br />dam at the top of the 15-mile reach <br />during 1991-1992. This relationship, <br />although variable, is remarkably con- <br />sistent from year to year; the upstream <br />areas inhabited by by larger adults are <br />consistently devoid of young-of-the- <br />year squawfish relative to the river <br />segment below Westwater Canyon and <br />the confluence with the Green River <br />(km 179-0) (Doug Osmundson, U.S. <br />Fish and Wildlife Service, Grand Junc- <br />tion, Colorado, unpublished data). <br />Kaeding and Zimmerman 1983; Kaeding et al. <br />1990; Karp and Tyus 1990; Valdez et al. 1990). <br />Squawfish and razorback sucker are almost always <br />captured in low or zero velocity habitats (Tyus 1984; <br />Osmundson and Kaeding 1989), which occur within <br />the active channel (e.g., eddies) or exist as backwa- <br />ters (e.g., back-bar channels) or floodplain wetlands <br />(i.e., flooded bottomlands) that are continually or <br />seasonally attached to the active channel. Squaw- <br />fish (Tyus 1991a, 1991b), and perhaps razorback <br />sucker (Minckley et al. 1991), must have access to <br />low velocity environments to mature. These obser- <br />vations strongly imply that low velocity habitats <br />are important feeding or resting areas or both, but <br />they do not imply that rivers of consistently low <br />velocity or volume are most suited to the endan- <br />gered fishes of the Upper Colorado River Basin. <br />Low velocity environments are formed and <br />maintained by complex hydrologic processes that