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created a trophic bottleneck for kokanee and the fishery collapsed in 1987 - 88. Since <br />1989, only incidental kokanee spawners have been observed in McDonald Creek and <br />the bald eagles have dispersed elsewhere (Spencer et al. 1991). <br />Accelerated Erosion <br />As in all river basins, soil and other mineral substratum are naturally eroded by <br />fluvial processes within the Flathead catchment. Owing to the porous nature of the <br />bedrock substrata and extensive tills of glacial origin, very little overland or sheet flow <br />occurs except during extreme precipitation events or during periods of intensely <br />accelerated snowmelt. Streams originate primarily as springbrooks fed by waters that <br />percolate into substrata from precipitation at higher altitudes. Springbrooks coalesce to <br />form the drainage network of the catchment. Therefore, most of the sediment loads <br />carried by the streams and rivers are derived from erosion of stream channels and <br />banks. <br />The rate of erosion is determined by channel morphology, slope, relative <br />erosiveness of streambank substrata and the intensity and duration of spates. Most of <br />the sediment load in the system is derived from Tertiary shales deposited as valley fill <br />and Quaternary tills and alluvium. These soils contain nitrogen and phosphorus that <br />are either contained within organic debris or associated with the clay lattice of the <br />mineral particles. Therefore, as much as 60% of the annual riverine nutrient load of the <br />Flathead River may be associated with sediment particles that are transported for short <br />time periods, most years during spring runoff, when the rivers and streams of the <br />catchment are flooding. Only about 10% of the nutrients associated with particles can <br />be assimilated by the biota (i.e., only about 10% of the particulate phosphorus is labile <br />or bioavailable, Ellis and Stanford 1986, 1988) and much of the load is either <br />deposited on the river floodplain or into the lakes as a short-term pulse event. In spite <br />of the low nutrient bioavai lability, the fertilization effect of the particulates eroded and <br />transported by fluvial processes is significant owing to 1) the oligotrophic nature of the <br />water bodies and 2) the dominance of the hydrograph and nutrient mass balance of <br />both rivers and lakes in the catchment by spring runoff. <br />Clearly, erosion is a natural process that both shapes the catchment landscape <br />and also to some extent fertilizes patches within the landscape. Natural (e.g., lightning- <br />caused fire, insect epidemics, beaver (Castor canadensis) and other large herbivore <br />influences) and human (e.g., road building, clearcutting) deforestation increases the <br />seasonal and annual variation in water yield, particularly during spring snowmelt <br />(Hauer and Blum 1991), thereby accelerating erosion of streambanks and increasing <br />sediment loads. Erosion of road surfaces and berms or stream crossings is of <br />14