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<br />Occasional Papers is published by Tulane University, Museum of Natural His- <br />tory. The Occasional Papers is dedicated primarily to Natural History: Sys- <br />tematics, Environmental Biology and Biogeography. The Occasional Papers will <br />appear irregularly in consecutively numbered issues. For information and policy <br />on exchanges, write to: Royal D. Suttkus, Director <br />Museum of Natural History <br />Tulane U niversi ty <br />Belle Chasse, Louisiana 70037 <br /> <br />When citing this publication, authors are requested to use the following: Occas. <br />Pap. Tulane U. Mus. Nat. Hist. <br /> <br />Cover photo: Mouth of the Little Colorado River in the Grand Canyon. <br /> <br />The National Fish and Wildlife Laboratory, Washington, D.C., supported pub- <br />lication of this number. <br /> <br />Printed by Drake Printers, Tuscaloosa, Alabama. <br />