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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />much of the upper basin still have a heightened spring runoff and low base flows (Van Steeter <br />k 1998). Water temperatures are similar to historic ranges and means (Kaeding and <br />son 1989), except for dam tailwaters that are released from deep, cold hypolimnetic <br />regions of reservoirs. Sediment transport has been modified in some regions of the upper basin, <br />particularly below dams (Andrews 1991). Invasion of alien plants and fish have also impacted <br />the historic ecosystem of the upper basin, resulting in extensive groves of nonnative tamarisk that <br />line and stabilize most of the river banks, sometimes at the exclusion of native willows and <br />cottonwoods. App ately 42 species of nonnative fishes compete with 13 maintem native <br />species in the uppe (Tyus et al. 1982), resulting in competitive and predatory forces that <br />ecline of some of the native species. These changes in hydrology, <br />temperature, sediment, and animal and plant diversity have resulted in subtle to extreme changes <br />in the riverine ecosystem. Extreme changes in macroinvertebrate and fish assemblages have <br />. taken place in reservoir basins and dam tailwaters (Holden and Crist 1979, Holden and Selby <br />1979), but further downstream, where flows are seasonally warmed and variable, much of the <br />historic macro invertebrate community i act (Haden 1998), although reduced or modified by <br />changes in flow regime and sediment smundson et al. 2000). Energy flow through the <br />ecosystem has also been altered by flow regulation and the presence of large nwnbers and <br />biomass of nonnative fishes (Osmundson et al. 2000). Despite these changes, there still remain <br />large, reproducing, and self-sustaining populations of many of the native species in the upper <br />basin. <br /> <br /> <br /> <br />In summary, the upper basin has about 1,355 km of ater riverine habitat suitable <br />for native warmwater fishes, and sustains reproducing, self-s 'ng populations of all native <br />fishes, except for bonytail and razorback sucker (some repro uction and possibly recruitment is <br />taking place with the razorback sucker). Despite reductions in flow volume and modifications to <br />flow regimes from dam operations and diversions, much of the historic structure and function of <br />the riverine ecosystem remain intact for several reaches of the upper basin. Because the upper <br />basin is now physically separated from the lower basin, the ecological features unique to the <br />upper reaches of the river continuwn have become isolated and these exert unique ec <br />forces on the evolutionary legacy of the native fishes, independent of the lower basin. <br /> <br /> <br />7 <br />