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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />INTRODUCTION <br /> <br />The endangered Colorado squawfish, Ptvchocheilus <br />lucius, is a large piscivorous cyprinid native to the <br />Colorado River Basin. Once abundant throughout the basin, <br />the species is now limited to the northern portion of its <br />historical range (Figure 1). The major causes of its <br />decline are generally attributed to the combined effects of <br />dam construction, introduction of non-native species, and <br />poor land use practices (Miller 1961; Minckley and Deacon <br />1968; Molles 1980; Behnke and Benson 1983; Carlson and Muth <br />1989). The small size of age-O Colorado squawfish entering <br />winter might be an important factor affecting recruitment to <br />the adult stock (Kaeding and Osmundson 1988). <br />In populations of temperate-zone fishes near the <br />northern limit of their range, the overwinter mortality of <br />age-O fish is often high (Oliver et ale 1979; Shuter and <br />Post 1989). This mortality may result from hypothermia, <br />predation, displacement by flooding, inadequate lipid <br />reserves leading to subsequent starvation, and physical <br />damage from anchor ice and frazil ice (Seelbach 1987). The <br />growing season shortens and the winter period lengthens as <br />one moves from south to north in the northern temperate <br />