<br />
<br />660
<br />
<br />COPEIA, 1989, NO.3
<br />
<br />trait. Sokal and Rohlf (1980) extrapolated fur-
<br />ther by stating that the trait was, in fact, irre-
<br />spective of variables such as age, sex, race, ex-
<br />perience, or even field of specialization.
<br />Individuals untrained in the complexities of
<br />taxonomic evaluation (to include children) did
<br />as well in phenetically grouping Caminalcules
<br />as did trained systematists.
<br />Does this mean that Rafinesque (1820) erred
<br />in the introductory pronouncement, which ex-
<br />tolled "seeing well" as "a high faculty of the
<br />mind"? We argue to the contrary. The "art of
<br />seeing well" (sensu Rafinesque) requires not
<br />necessarily a "high faculty of the mind," but
<br />instead innately human traits like perseverence
<br />and discipline. All humans apparently have the
<br />capability to group organisms and objects using
<br />shape and size parameters, but many would elect
<br />not to do so based upon a personal lack of per-
<br />severence and diligence required by such tasks.
<br />These factors may be critical in promoting the
<br />trained eye of a systematist.
<br />Results of our investigations impinge heavily
<br />on the complex program to preserve and man-
<br />age endangered fishes in the Colorado River
<br />(USFWS, 1987c), a program that involves state,
<br />federal, private, and academic groups. Deep,
<br />emotionally-charged issues which sustain and
<br />occasionally divide these groups range from
<br />water-rights acquisition to field identification of
<br />species. With regard to the last issue, our results
<br />indicate that qualitative features of certain phe-
<br />notypic regions are adequate to distinguish at
<br />least two different species, G. robusta and the
<br />endangered G. cypha, occurring syntopically in
<br />the Yampa River. Our investigation unequiv-
<br />ocally demonstrates that this relatively undis-
<br />turbed reach of stream is important habitat for
<br />G. 0pha. Efforts in 1987 produced 8.9% G. cypha
<br />(32 of 358 total specimens of Gila collected), in
<br />a variety of sizes and of both sexes in repro-
<br />ductive condition. Unfortunately, no G. elegans
<br />was collected, so the use of qualitative charac-
<br />ters could not be tested on a multi-species group,
<br />nor are we able to comment on status of bony tail
<br />in the study area other than to further attest to
<br />its rarity.
<br />Identification of G. cypha and its absolute and
<br />relative abundance serve to demonstrate that
<br />the lower Yampa River represents a relatively
<br />pristine example of what was once the entire
<br />Colorado River basin (Fradkin, 1984). As such,
<br />it may well represent one of the last opportu-
<br />nities for the system's native fishes to survive,
<br />
<br />as also indicated by the presence of well docu-
<br />mented spawning grounds for the endangered
<br />Colorado squawfish (Pt)'chocheilus lucius) (Tyus,
<br />1986), occurrences of razorback suckers (X)'-
<br />rauchen texanus) that are otherwise exceedingly
<br />rare in the upper Colorado basin (McAda and
<br />Wydoski, 1980; Tyus, 1987), as well as a possible
<br />refuge for the endangered bony tail (Valdez and
<br />Clemmer, 1982). Every effort should be made
<br />to retain the Yampa River as a critical refugium
<br />for G. c)'pha and other Colorado River fishes,
<br />and as an area for further study of these unique
<br />and imperiled species.
<br />
<br />ACKNOWLEDGMENTS
<br />
<br />P. Clevenger, C. Karp, P. Minckley, K. Pau-
<br />lin, L. Trinca, and C. Williams collected most
<br />of the data for this project. Their diligence and
<br />industry are to be commended. G. R. Smith
<br />provided early discussions toward development
<br />of ranking factors for this group of fishes. P. C.
<br />Marsh and G. R. Smith read and improved the
<br />paper. Funding and logistic support was pro-
<br />vided by Arizona State University, U.S. Bureau
<br />of Reclamation, and U.S. Fish and Wildlife Ser-
<br />vice. Issuance of permits by the States of Col-
<br />orado and Utah, U.S. National Park Service
<br />(Dinosaur National Monument), and U.S. Fish
<br />and Wildlife Service is gratefully acknowl-
<br />edged.
<br />
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