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<br />Backwaters <br /> <br />One of the more dramatic relationships seen in this analysis was <br />between discharge and area of backwaters (Figure 7). Two thresholds <br />occurred, one subtle and one more dramatic. At low flows, the surface area <br />of backwaters was low and represented by small discrete units formed by <br />shoreline depressions (Figure 3). At about 6,000 cfs, these depressions <br />were inundated by flow-through and many of these backwaters were lost as <br />others developed (Figure 5). However, when river discharge reached about <br />10,000 cfs, a nearly 50-fold increase in backwater area was seen. This <br />occurred during runoff when the river flooded adjacent low-lying depressions <br />1 i ke 01 d si de channel s and man-made features such as gravel pi ts. <br />Backwaters were less numerous at this time than during low flow, but each <br />individual backwater encompassed a much larger area. <br /> <br />The occurrence of backwaters appears to coincide with certain events in <br />the 1 ife history of the river fishes. Spawning for many of the riverine <br />species in this area occurs during the descending limb of runoff and these <br />large backwaters may serve as areas for resting, maturation, and spawning as <br />flows decreased, and backwaters become more numerous but smaller in area. <br />The young fish occupied these small, warm, sheltered areas to find food and <br />escape deep-water predators. It was a1 so noted that these large backwaters <br />were usually warmer than the main channel, by 1-50C. This may enable <br />certain species to accumulate temperature degree-days needed for <br />reproductive maturation. <br /> <br />Flooded Woodlands <br /> <br />The area of flooded woodlands, like that of backwaters, also increased <br />dramatically at about 10,000 cfs discharge (Figure 7). This habitat <br />consisted mostly of flooded willows and other riparian vegetation that was <br />surrounded by water when the river exceeded its common-flow banks. This <br />habitat appears to be important to small fish as feeding areas and as <br />shelter from the main channel turbulence during runoff. <br /> <br />Embayments and Concavities <br /> <br />The areas of embayments and concavities (Fi gure 7) did not bear a <br />predictable relationship to discharge. These small habitats depend on <br />shore1 ine depressions which apparently occur randomly and at all el evations <br />within the floodplain. Embayments usually occur more frequently in sandy <br />areas where shi fting of the substrate is more common than in rocky areas <br />such as the study area (Valdez and Wick 1981). Larval fish surveys in <br />Colorado during 1979-1981 indicated that shorel ine embayments and <br />concavities are an important habitat feature to larval Colorado squawfish <br />(Haynes et a1. 1984). <br /> <br />Rapids, Runs and Riffles <br /> <br />The occurrence and areas of rapids and runs (Fi gure 8) seemed to follow <br />a similar pattern. However, rapids occurred only above 10,000 cfs, when the <br />ri ver turbu1 ence was severe, enough to create these areas. The area of <br />riffles (Figure 8) occurred in inverse relationship to those of rapids and <br />runs. Runs and riffles appear to be important habitats for suckers and dace <br />as feeding areas and perhaps as spawning areas (Valdez et al. 1982). <br />However, few fish were found in rapids. <br /> <br />257 <br />