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<br />Eddies <br /> <br />The occurrence and surface area of eddies varied directly with <br />discharge (Figure 8), and appeared to be a function of river hydraulics. <br />Eddies provide a food source and habitat for many chubs and suckers. <br />Roundtail chubs are often abundant along "eddy fences", or interfaces <br />between main current and backflow of eddies where they feed on insects <br />drifting in the main channel that are swept into eddies (Valdez et al. <br />1982) . <br /> <br />Shorelines and Rubble Flats <br />--- <br /> <br />Shorel ine habitat did not appear to vary with discharge (Figure 9). <br />Although never a constant area, dramatic changes in this habitat did not <br />occur with flow and stage changes. Shoreline habitats ~nd associated <br />features often support the majority of riverine fish populations (Hynes <br />1970). Their consistent area and physical nature (lack of turbulence <br />associated with midchannel habitats) may explain their importance as fish <br />habitats. Rubbl e flats occurred only at flows between about 5,000 and <br />12.000 cfs (Figure 9). This habitat appears to be important to some species <br />of fish that associate with shallow riffles, such as speckled dace and young <br />suckers. <br /> <br />Isolated Pools <br /> <br />Isolated pools occurred at most flow stages (Figure 9), as a result of <br />low-lying areas losing contact with the river as discharge decreased. In <br />three field seasons of seining in the Colorado, Gunnison and Dolores Rivers, <br />Valdez et al. (1982) found many non-native species but few native fishes <br />stranded and dying in isolated pool s. Apparently, the native fishes are <br />keenly aware of changing flow stages and leave backwaters and side channels <br />before they become isolated pools. <br /> <br />Pools <br /> <br />The trace of the thalweg in March revealed a single pool area about 15 <br />feet deep immediately downstream from the Battlement Mesa Bridge. As <br />discharge increased, velocity through this area increased changing the pool <br />to a deep run. Pool-like character was observed for this area only at the <br />lowest flows (Table 1). <br /> <br />It is currently bel ieved that Colorado squawfish spawn in riffles at <br />the upper end of resting pOOls (Archer and Tyus 1984) in much the same <br />manner as the northern squawfi sh of the Co 1 umbi a Ri ver (Beamsderfer and <br />Congleton 1981). Seethaler (1978) first hypothesized this idea and it was <br />later tested by Tyus et al. (1982) and Lamarra et al. (1983) in the Yampa <br />River. Recent radiotelemetry studies strongly suggest the use of this <br />habitat type for staging and resting of squawfish during spawning activities <br />(Archer and Tyus 1984). <br /> <br />Habitat Dynamics <br /> <br />A synthesis of habitat areas over time (Figure 10) and by discharge <br />(Figure 11), clearly shows the total habitat:discharge relationship. During <br />the period 6 June through 2 September, total habitat area was at a maximum <br />during peak flow in late June and declined to a minimum level in late July <br />through early September. <br /> <br />260 <br />