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<br />(heterotrophy) and low photosynthesis (autotrophy). <br />However, if autotrophy (high primary production) is <br />greater than heterotrophy, then the ratio will be greater <br />than 1. Thus, continuous monitoring of production and <br />respiration rates through the seasons will allow a direct <br />determination of the balance between primary production <br />and consumption of organic material. It is this balance <br />which will be a measure of the relative health of the <br />system from which departures may be identified and related <br />to the system structural components. This mechanism of <br />classifying the White River system is of great importance <br />in determinimg potential structural or functional changes <br />in the biotic component in a timely manner. <br /> <br />RATIONALE OF THE ECOSYSTEM APPROACH <br /> <br />Ecosystem resolution can be carried from the most <br />gross features (macrostructure/function) to the most <br />miniscule (microstructure/function). What must be kept in <br />mind with the management/monitoring program, therefore is <br />a reasonable perspective based on logic, economics, <br />regulatory requirements, ecosystem knowledge and the <br />expected pathway or mechanism of effects. <br /> <br />Based on the previous view of the aquatic ecosystem, <br />we can begin, for example, at the macro-level by <br />establishing relationships between primary production and <br />environmental physical factors such as light, temperature, <br />water chemistry and so forth. For the invertebrate or <br />consumer populations we can establish relationships <br />between environmental factors, terrestrial organic input, <br />primary production, and production of the consumers. At <br />the decomposer level we can measure indices of microbial <br />community dynamics such as enzymatic activities coupled to <br />community metabolism and relate this to stream physical <br />environment, organic matter and water chemistry. <br />Overlaying all these subsystems we can determine <br />production/respiration (P/R) ?nd decomposition rates as <br />related to environmental factors through the year to <br />arrive at a temporal, quantitative picture of gross <br />community behavior. <br /> <br />By considering an ecosystem in terms of energy or <br />carbon flow, measuring production and consumption between <br />each subsystem and for the overall system, and calculating <br />mass balances, we will arrive at a quantitative <br />description of the ecosystem. This quantitative <br />description then becomes a statistically based valid <br />reference with which to compare future system behavior for <br />determination of departures from normality. If the <br />relationships under consideration are valid, departures <br /> <br />281 <br />