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<br />SUBADULT HUMPBACK CHUB IN THE COLORADO RIVER <br /> <br />275 <br /> <br />Table VI. Results of generalized, randomized block MANOV A for mean depth, veloc- <br />ity and cover among shoreline types and reaches (for reaches I and 2 only). DF, degrees <br />of freedom <br /> <br />Source <br /> <br />DF <br /> <br />F <br /> <br />Shoreline (S) <br />Reach (R) <br />RxS <br /> <br />15, 152 <br />3,55 <br />15,152 <br /> <br />Wilks' ). <br /> <br />P <br /> <br /><0.001 <br /><0.001 <br />0.003 <br /> <br />0.198 <br />0.611 <br />0.554 <br /> <br />8.09 <br />11.68 <br />2.42 <br /> <br />Relationships between subadult humphack chub and geomorphology <br /> <br />Fish densities did not monotonically decline from upstream to downstream or show any other <br />recognizable pattern; however, mean relative density varied substantially among shoreline types (Figure <br />4). In addition, other species, such as speckled dace, not reported here, differed in habitat use and relative <br />abundance from that of subadult humpback suggesting that sampling efficiency bias among shoreline <br />types was minimal (Valdez and Ryel, 1995). These results imply that subadults were quickly dispersing <br />and then preferentially using specific shoreline types along the river corridor while avoiding others. <br />Discriminant functions analysis showed that subadult humpback chub used locations that were physically <br />different than unoccupied areas (Table VII). Areas with fish had more cover (p < 0.001) and lower <br />velocity (p = 0.10) than those without but did not differ significantly in depth (p = 0.84). <br />Estimates of overall fish abundance, based on shoreline availability and shoreline use, were highest in <br />reach 3 (82 fish caught per 10 h of fishing), intermediate in reach I (50 fish caught per 10 h fishing) and <br />lowest in reach 2 (33 fish caught per 10 h fishing). However, a GRB ANOV A showed a significant <br />interaction between reach and shoreline type, suggesting that the pattern of shoreline selection varied <br />between reaches (Table VIII). <br />Overall, densities were highest in vegetated shorelines, followed by talus and debris fan shorelines. <br />Bedrock, cobble and sand shorelines had low densities of subadults. However, when considering the <br />interaction with reach, relative densities in bedrock shorelines in reach 2 were high compared with <br />bedrock shorelines in reaches I and 3, whereas relative densities in talus shorelines in reach 2 were low <br /> <br />: <br /> <br />3 <br />2 <br /> <br />Depth (m) <br /> <br /> <br />o <br />0.60 <br />Velocity (m/s) <br />0.45 <br />0.30 <br />0.15 <br />0.0 <br />100 <br />80 <br />60 <br />40 <br />20 <br />o <br /> <br /> <br />Cover (%) Reach 1 <br />. Reach 2 <br /> <br /> <br />BE CB OF SA TS VG <br />Shoreline type <br /> <br />Figure 3. Physical differences in depth, velocity and cover among shoreline types and between Reaches I and 2. Be, bedrock; CB, <br />cobble; DF, debris fan; SA, sand; TS, talus; and VG, vegetation <br /> <br /><<:J 1998 John Wiley & Sons, Ltd. <br /> <br />Regul. Rivers: Res. Mf(mt. 14: 267-284 (1998) <br />