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<br />SUBADULT HUMPBACK CHUB IN THE COLORADO RIVER <br /> <br />269 <br /> <br />. <br />.. <br /> <br />Historically, the Colorado River was a highly turbid, highly fluctuating system. Temperature varied <br />seasonally from near freezing to almost 300C. Flows could range annually from approximately 60 m3 s - I <br />during winter months to over 3000 m3 s - I (US Department of Interior, Bureau of Reclamation, 1995). <br />The dam has altered the hydrology of the river by eliminating large annual flood events, maintaining <br />artificially high base flows, trapping sediment and altering physio-chemical conditions (Stanford and <br />Ward, 1991). Water discharged from the dam is approximately 7-11oC and warms an average of 10 every <br />50 km downstream in the summer (Valdez and Ryel, 1995). <br />From the period of dam closing in 1962-1991, discharge fluctuated within a range of 60-850 m3 s - I <br />with unrestricted ramping rates. To reduce these fluctuations, the USBR implemented interim flows in <br />August 1991. This operating regime requires a minimum of 225 m3 s -I and a maximum of 550 m3 s - 1. <br />Up ramping (rate of discharge increase) does not exceed 70 m3 s -1 h -1, and down-ramping (rate of <br />discharge decrease) does not exceed 45 m3 s - I h - 1 (US Department of Interior, Bureau of Reclamation, <br />1995). <br />Within the Grand Canyon, the study area begins at the confluence of the Colorado River with the LCR <br />(RM 61.4), a calcium carbonate-enriched, spring-fed tributary to the Colorado River in Grand Canyon <br />(Figure I). The LCR flows clear except during heavy, upland rainstorms when it floods and becomes <br />sediment laden. With a base flow of 5.5 m3 s - I and a lO-year recurrence interval flood of 425 m3 s - 1, <br />the LCR is the largest tributary entering the Colorado River in Grand Canyon. <br /> <br />Geomorphology <br /> <br />Grand Canyon is the result of concurrent uplifting of the Colorado Plateau and down cutting of the <br />Colorado River through more than 1500 m of sedimentary and metamorphic rock (Beus and Morales, <br />1990). The river through Grand Canyon drops over 500 m in 450 km (279 miles) as a series of long, flat <br />stretches interrupted by steep drops. Half of the drop in elevation is in rapids, which constitute only 9% <br />of the total distance. Most of the rapids are formed by debris fans at the mouths of ephemeral tributaries <br />that deposit poorly sorted debris ranging in size from huge boulders to sand. These debris fans occur <br />where local fractures or faults transect the canyon at the river (Dolan and Howard, 1978). Debris flows <br />and tributaries deposit enough sediment to constrict the river, forming rapids. Associated with rapids are <br />large downstream zones of recirculation known as eddy complexes (Schmidt and Graf, 1990). <br />Channel slope and morphology in Grand Canyon change along the river corridor with shifts in local <br />geology and tributary influence (Leopold, 1969; Kieffer, 1990). More resistant lithology creates a narrow, <br /> <br />I <br /> <br /> <br />NEVADA: <br />1 <br />I. _ _ _ ~T:\!:l_ _ _ _ _ _ _ _ _ <br />: ARIZONA <br /> <br />, <br />I <br />I <br />I <br /> <br />IN <br /> <br />Study Area <br /> <br />Figure 1. Map of the Colorado River through Grand Canyon <br /> <br />~ 1998 John Wiley & Sons, Ltd. <br /> <br />Rel(ul. Rivers: Res. Ml(mt. 14: 267-284 (1998) <br />