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<br />282 <br /> <br />Y.K. CONVERSE ET AL. <br /> <br />These thermal constraints together with a decrease in habitat quality due to artificially high base flows <br />and a change in food availability have probably inflicted an extreme and detrimental post-dam environ- <br />mental change on subadult humpback chub in the Colorado River through Grand Canyon that limits <br />their growth and survival. Yet, to date, the effect of artificially elevated base flows has not been <br />adequately included in the evaluation of Glen Canyon Dam operations on native fish habitat in the <br />Colorado River through Grand Canyon. <br /> <br />ACKNOWLEDGEMENTS <br /> <br />This research was supported by the United States Bureau of Reclamation through a contract to BiojWest, <br />Inc. We thank the many BiojWest and Bureau of Reclamation employees who assisted in data collection, <br />especially Bill Masslich, Leslie Brown, Peter Weiss and Bryan Cowdell. Todd Crowl, Mike O'Neill and <br />Lou Pisacane provided constructive criticisms throughout the study. We thank Jack Schmidt for sharing <br />ideas during early phases of the study and providing raw channel width and depth data. <br /> <br />REFERENCES <br /> <br />Angradi, T.R. 1994. 'Trophic linkages in the lower Colorado River: multiple stable isotope evidence', J. North Am. Benthol. Soc., <br />13, 479-495. <br />Arizona Game and Fish Department 1994. Glen Canyon Environmental Studies Phase 1/ 1993 Annual Report. Prepared for the <br />Bureau of Reclamation, Upper Colorado Region, Glen Canyon Environmental Studies. Flagstaff, AZ. Cooperative Agreement <br />No. 9-FC-40-07940. Arizona Game and Fish Department, Phoenix. 43 pp. <br />Belknap, B. and Evans, L.B. 1989. Belknap's Grand Canyon River Guide. Westwater Books, Denver, CO. 44 pp. <br />Berry, C.R. and Pimentel, R. 1985. 'Swimming performances of three rare Colorado River fishes', Trans. Am. Fish. 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