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<br />SWIMMING PERFORMANCES OF RARE COLORADO RIVER FISHES <br /> <br />401 <br /> <br />. <br /> <br />Zimmer and Broughton 1965). Our data should <br />partially define one of a dozen biological criteria <br />needed to design an experimental fish ladder <br />(Truebe and Drooker 1982). Additionally, data <br />such as ours are required in formulas used to <br />design culverts for fish passage (Watts 1974; Ev- <br />ans and Johnston 1980). <br />We attempted to simulate conditions that these <br />fish might encounter after hydrologic alterations <br />in stream habitat; however, our data are incom- <br />plete for most engineering and construction plan- <br />ning. Many other aspects of the swimming per- <br />formance and behavior of each life stage must <br />be examined before developers can adequately <br />consider fish preferences. Our data are probably <br />conservative because of the limitations ofthe test <br />apparatus and the use of unexercised fish. The <br />effect of fish length, water velocity, and water <br />temperature on swimming ability could have been <br />predicted from existing literature (Beamish 1978). <br />Unique to our study was the opportunity to eval- <br />uate the specific abilities of these rare fishes and <br />the finding that their abilities were similar to <br />those of other fishes of similar size. This is en- <br />couraging because existing data for other species <br />can augment our data in solving fish-passage and <br />entrainment problems. <br /> <br />J <br /> <br />Acknowledgments <br />Funds for this work were provided by the U.S. <br />Fish and Wildlife Service Colorado River Fish- <br />ery Project and the Bureau of Reclamation. R. <br />Bulkley, L. Kaeding, and H. Tyus reviewed an <br />early draft of this paper; A. Burgess and J. Kehoe <br />conducted most of the swimming tests. This <br />project was administered by the Utah Cooper- <br />ative Fishery Research Unit, which is jointly <br />sponsored by the U.S. Fish and Wildlife Service, <br />the Utah Division of Wildlife Resources, and <br />Utah State University. <br /> <br />References <br /> <br />ADAMS, B. D., AND V. A. LAMARRA, editors. 1983. <br />Aquatic resources management of the Colorado <br />River ecosystem. Ann Arbor Science, Ann Arbor, <br />MI, USA. <br />BARNES, D. 1976. Development document for best <br />technology available for the location, design, con- <br />struction, and capacity of cooling water intake <br />structures for minimizing adverse environmental <br />impact. U.S. Environmental Protection Agency <br />4401l76/015a, Washington, DC, USA. <br />BEAMISH, F. W. H. 1970. Oxygen consumption of <br />largemouth bass, Micropterus salmoides, in rela- <br /> <br />tion to swimming speed and temperature. Cana- <br />dian Journal of Zoology 49:1221-1228. <br />BEAMISH, F. W. H. 1978. 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