<br />Clearly, recent research by CRFP, DOW, and others
<br />has yielded important information, however, we
<br />must be willing to admit that our knowledge is
<br />largely surficial and that questions such as
<br />those above must be answered if this endemic fish
<br />is to be effectively managed.
<br />
<br />Perhaps the most crucial research/management
<br />implication of past studies is that a coordinated
<br />research effort throughout the upper basin, based
<br />upon a critically organized and goal-oriented
<br />research design, will yield needed empirical data
<br />in the shortest possible time. It is important,
<br />however, that a realistic time frame be allocated
<br />for this work--researchers who have been involved
<br />with implementing past field studies and inter-
<br />preting collected data realize all too well the
<br />difficulties of deriving meaningful information
<br />on this elusive species.
<br />
<br />The questions and comments offered in the above
<br />synthesis have obvious broad applicability in
<br />virtually all circumstances where the recovery
<br />and management of an endangered species is an
<br />issue. Important and useful insights relative to
<br />the ecological requirements of the Colorado
<br />squawfish may be derived from knowledge of the
<br />needs of large wide-ranging predators in general,
<br />and more speciiically, from research conducted
<br />elsewhere on large potamodromous cyprinids.
<br />Unquestionably, habitat alterations resulting
<br />from water resource development in the upper
<br />Colorado River basin will continue as states
<br />endeavor to develop their full entitlements under
<br />the 1922 Colorado River Compact and the 1948
<br />Upper Colorado River Basin Compact (Bishop and
<br />Porcella 1976). These compacts insure that the
<br />upper basin deliver specified flows to lower
<br />basin states and that Utah is entitled to minimal
<br />flows from Colorado. Accordingly, the water
<br />remaining in upper basin river channels after all
<br />entitlements are met will represent the totality
<br />of potential habitat available for the Colorado
<br />squawfish and other rare Colorado River endemic
<br />fishes. If remaining flows are to provide
<br />acceptable fish habitat, they must be budgeted to
<br />provide for the species' life requisites in terms
<br />of timing, quantity, and quality. Inferences
<br />made from broadly applicable ecological knowledge
<br />can provide important guidelines for flow
<br />budgeting considerations until more species-
<br />specific information has been derived. Clearly,
<br />the highly regulated Colorado River offers
<br />continually diminishing unaltered habitat for the
<br />perpetuation of a species that evolved under
<br />vastly different ecological conditions from those
<br />which it now faces. Yampa Canyon in northwestern
<br />Colorado is one of the few remaining river
<br />reaches in the upper basin that has retained much
<br />of its pre-development (Le., "wilderness")
<br />character and provides an excellent outdoor
<br />laboratory for investigating cause and effect
<br />relationships for this particular endangered
<br />species. In a general sense, knowledge gained to
<br />date from studies in Yampa Canyon is illustrative
<br />of the connection between the preservation of
<br />rare species and the maintenance of large
<br />wilderness areas. In the cases of the Colorado
<br />squawfish, its future throughout much of
<br />
<br />its existing range appenrs to be closely tied to
<br />the continuation of practices that insure the
<br />natural features of Yampa Canyon.
<br />
<br />REFERENCES
<br />
<br />Archer, D.L., H.M. Tyus, and R.A. Valdez. Field
<br />methodologies of the Fish and Wildlife
<br />Service's Colorado River Fisheries Project.
<br />Transactions of the Bonneville Chapter,
<br />American Fisheries Society, Vol. 1980: 13-36.
<br />
<br />Banks, J.L. Fish species distribution in
<br />Dinosaur National Monument during 1961 and
<br />1962. Fort Collins, co: Colorado State
<br />University; 1964. 96 p. M.S. thesis.
<br />
<br />Bishop, A.B., and D.B. Porcella. Physical and
<br />ecological aspects of the upper Colorado River
<br />basin in relation to energy development and
<br />environmental problems. Utah Water Research
<br />Laboratory, Utah State University; 1976. 32 p.
<br />
<br />Colorado River Fishes Recovery Team.
<br />squawfish recovery plan. U.S. Fish
<br />Service. 1978. 65 p.
<br />
<br />Colorado
<br />and Wildlife
<br />
<br />Ehrenfeld, D.W. The conservation of non-
<br />resources. American Scientist. 65:648-656;
<br />1976.
<br />
<br />Hamman, R.L. Spawning and culture of Colorado
<br />squawfish in raceways. Progressive Fish-
<br />Culturist. 43: 173-177; 1981.
<br />
<br />Haynes, C.M., and R.T. Muth. Reproduction of
<br />Colorado squawfish, Ptychocheilus lucius, in
<br />the Yampa River, Colorado (1980-83). Paper
<br />presented at 15th Annual Symposium, Desert
<br />Fishes Council, Death Valley National Monument,
<br />CA. 17-19 Nov., 1983 (in press).
<br />
<br />Haynes, C.M., and R.T. Muth. Identification of
<br />habitat requirements and limiting factors for
<br />Colorado squawfish and humpback chubs. In:
<br />Wildlife Research Report, part 2, Nongame
<br />Investigations; Jan. 1984. Colorado Division of
<br />Wildlife, Fort Collins, CO: 177-202.
<br />
<br />Haynes, C.M., and R.T. Muth. Downstream
<br />transport of larval Colorado squawfish in the
<br />Yampa River, Colorado (U.S.A.). (in
<br />preparation).
<br />
<br />Haynes, C.M., T.A. Lytle, E.J. Wick, and R.T.
<br />Muth. Larval Colorado squawfish (Ptychocheilus
<br />lucius Girard) in the upper Colorado River
<br />basin, Colorado, 1979-1981. The Southwest
<br />Naturalist 29(1):21-33; 1984.
<br />
<br />Holden, P.B. Distribution, abundance, and life
<br />history of the fishes of the upper Colorado
<br />River basin. Logan, UT, Utah State University;
<br />1973. 59 p. Ph.D. dissertation.
<br />
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