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<br />Clearly, recent research by CRFP, DOW, and others <br />has yielded important information, however, we <br />must be willing to admit that our knowledge is <br />largely surficial and that questions such as <br />those above must be answered if this endemic fish <br />is to be effectively managed. <br /> <br />Perhaps the most crucial research/management <br />implication of past studies is that a coordinated <br />research effort throughout the upper basin, based <br />upon a critically organized and goal-oriented <br />research design, will yield needed empirical data <br />in the shortest possible time. It is important, <br />however, that a realistic time frame be allocated <br />for this work--researchers who have been involved <br />with implementing past field studies and inter- <br />preting collected data realize all too well the <br />difficulties of deriving meaningful information <br />on this elusive species. <br /> <br />The questions and comments offered in the above <br />synthesis have obvious broad applicability in <br />virtually all circumstances where the recovery <br />and management of an endangered species is an <br />issue. Important and useful insights relative to <br />the ecological requirements of the Colorado <br />squawfish may be derived from knowledge of the <br />needs of large wide-ranging predators in general, <br />and more speciiically, from research conducted <br />elsewhere on large potamodromous cyprinids. <br />Unquestionably, habitat alterations resulting <br />from water resource development in the upper <br />Colorado River basin will continue as states <br />endeavor to develop their full entitlements under <br />the 1922 Colorado River Compact and the 1948 <br />Upper Colorado River Basin Compact (Bishop and <br />Porcella 1976). These compacts insure that the <br />upper basin deliver specified flows to lower <br />basin states and that Utah is entitled to minimal <br />flows from Colorado. Accordingly, the water <br />remaining in upper basin river channels after all <br />entitlements are met will represent the totality <br />of potential habitat available for the Colorado <br />squawfish and other rare Colorado River endemic <br />fishes. If remaining flows are to provide <br />acceptable fish habitat, they must be budgeted to <br />provide for the species' life requisites in terms <br />of timing, quantity, and quality. Inferences <br />made from broadly applicable ecological knowledge <br />can provide important guidelines for flow <br />budgeting considerations until more species- <br />specific information has been derived. Clearly, <br />the highly regulated Colorado River offers <br />continually diminishing unaltered habitat for the <br />perpetuation of a species that evolved under <br />vastly different ecological conditions from those <br />which it now faces. Yampa Canyon in northwestern <br />Colorado is one of the few remaining river <br />reaches in the upper basin that has retained much <br />of its pre-development (Le., "wilderness") <br />character and provides an excellent outdoor <br />laboratory for investigating cause and effect <br />relationships for this particular endangered <br />species. In a general sense, knowledge gained to <br />date from studies in Yampa Canyon is illustrative <br />of the connection between the preservation of <br />rare species and the maintenance of large <br />wilderness areas. In the cases of the Colorado <br />squawfish, its future throughout much of <br /> <br />its existing range appenrs to be closely tied to <br />the continuation of practices that insure the <br />natural features of Yampa Canyon. <br /> <br />REFERENCES <br /> <br />Archer, D.L., H.M. Tyus, and R.A. Valdez. Field <br />methodologies of the Fish and Wildlife <br />Service's Colorado River Fisheries Project. <br />Transactions of the Bonneville Chapter, <br />American Fisheries Society, Vol. 1980: 13-36. <br /> <br />Banks, J.L. Fish species distribution in <br />Dinosaur National Monument during 1961 and <br />1962. Fort Collins, co: Colorado State <br />University; 1964. 96 p. M.S. thesis. <br /> <br />Bishop, A.B., and D.B. Porcella. Physical and <br />ecological aspects of the upper Colorado River <br />basin in relation to energy development and <br />environmental problems. Utah Water Research <br />Laboratory, Utah State University; 1976. 32 p. <br /> <br />Colorado River Fishes Recovery Team. <br />squawfish recovery plan. U.S. Fish <br />Service. 1978. 65 p. <br /> <br />Colorado <br />and Wildlife <br /> <br />Ehrenfeld, D.W. The conservation of non- <br />resources. American Scientist. 65:648-656; <br />1976. <br /> <br />Hamman, R.L. Spawning and culture of Colorado <br />squawfish in raceways. Progressive Fish- <br />Culturist. 43: 173-177; 1981. <br /> <br />Haynes, C.M., and R.T. Muth. Reproduction of <br />Colorado squawfish, Ptychocheilus lucius, in <br />the Yampa River, Colorado (1980-83). Paper <br />presented at 15th Annual Symposium, Desert <br />Fishes Council, Death Valley National Monument, <br />CA. 17-19 Nov., 1983 (in press). <br /> <br />Haynes, C.M., and R.T. Muth. Identification of <br />habitat requirements and limiting factors for <br />Colorado squawfish and humpback chubs. In: <br />Wildlife Research Report, part 2, Nongame <br />Investigations; Jan. 1984. Colorado Division of <br />Wildlife, Fort Collins, CO: 177-202. <br /> <br />Haynes, C.M., and R.T. Muth. Downstream <br />transport of larval Colorado squawfish in the <br />Yampa River, Colorado (U.S.A.). (in <br />preparation). <br /> <br />Haynes, C.M., T.A. Lytle, E.J. Wick, and R.T. <br />Muth. Larval Colorado squawfish (Ptychocheilus <br />lucius Girard) in the upper Colorado River <br />basin, Colorado, 1979-1981. The Southwest <br />Naturalist 29(1):21-33; 1984. <br /> <br />Holden, P.B. Distribution, abundance, and life <br />history of the fishes of the upper Colorado <br />River basin. Logan, UT, Utah State University; <br />1973. 59 p. Ph.D. dissertation. <br /> <br />195 <br />