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<br />. . <br /> <br />a <br /> <br />HOLDEN AND STALNAKER-COLORADO RIVER FISHES <br /> <br />219 <br /> <br />Many of the streams in the upper basin <br />flow free, but two major reservoirs were <br />built in the 1960's: Baming Gorge Reser- <br />voir located on the upper Green River and <br />Lake Powell just upriver from Marble and <br />Grand Canyons (Fig. 1). <br />Evermann and Rutter (1895) summarized <br />collections from the Colorado River made <br />prior to 1894. Taba, Murphy, and Frost <br />(1965) made collections in the Colorado <br />River for several miles below Moab, Utah, <br />and Smith (1959) and McDonald and Dot- <br />son (1960) conducted preimpoundment sur- <br />veys in Glen Canyon. A fish eradication <br />program in September 1962 (Binns et al. <br />1964) effectively eliminated most native <br />species above and for many kilometers <br />below Flaming Gorge Dam before closure <br />in November 1962. Vanicek, Kramer, and <br />Franklin (1970) documented the distribution <br />of native fishes below this dam following <br />closure, and summarized preimpoundment <br />surveys. They found no reproduction of na- <br />tive fishes in the cold tail waters of the <br />reservoir for 104 kilometers downstream. <br />At that point, the Yampa River joins the <br />Green and ameliorates the coldness of the <br />Green somewhat. Areas noi previously <br />studied include the lower 400 km of the <br />Green River, the Yampa River, and the <br />Colorado River above Moab, Utah. <br />Minckley and Deacon (1968) reported <br />two squawfish taken in the Grand Canyon <br />in the mid 1960's. No subsequent accounts <br />of squawfish from this canyon have been <br />published. Minckley (1973) reported the <br />humpback sucker as nearing extinction <br />helow Lake Mohave. <br />Presently, Colorado squawfish and <br />humpback chub are considered "En- <br />dangered" by the U.S. Department of the <br />Interior (1973). Humpback sucker and <br />bony tail chub are also rare (Miller 1972). <br />The present study was undertaken to de- <br />termine the status of fishes in the middle <br />and upper Colorado River basins, with <br />special emphasis on endemic, large-river <br />forms. <br />The objectives were: (1) to determine <br />distribution and relative abundance of <br />fishes in the Colorado River basin above <br /> <br />Lake Mead, and (2) to identify areas im- <br />portant for reproduction of endangered <br />forms. <br /> <br />STUDY AREA <br /> <br />That part of the Colorado River basin <br />(Fig. 1) considered in this study includes <br />the Green River system from Craig, Col- <br />orado; the Yampa River to the junction of <br />the Green and Colorado Rivers; and the <br />Colorado River from Hifle, Colorado to <br />Lake Mead, excluding Lake Powell. Sev- <br />eral larger tributaries also included were <br />the Gunnison River below Delta, Colorado, <br />and the Dolores, Price, Whitc, and Little <br />Snake Rivers. <br />The main system was divided into 12 <br />sampling areas: <br />1. The Upper Yampa River from Craig, <br />Colorado, to Juniper Springs, Colorado, <br />approximately 80 km. <br />2. The Lower Yampa Hiver in Yampa <br />Canyon, Dinosaur National Monument, ap- <br />proximately 72 km. <br />3. The Green River from the mouth of <br />the Yampa River to the south boundary of <br />Dinosaur National Monument, about 32 <br />km. <br />4. The Green River from Ouray to <br />Green River, Utah, including Desolalion <br />and Grey Canyons, about 192 km. <br />5. The Green River from Mineral Bot- <br />tom to its mouth, most of which is in <br />Canyonlands National Park, about 104 km. <br />6. The Colorado River for 16 km up- <br />stream from mouth of the Green River. <br />7. The Colorado River upstream from <br />Moab, Utah, approximately 32 km. <br />8. The Colorado River from Grand Junc- <br />tion to Fruita, Colorado, about 16 km. <br />9. The Colorado River near Rifle, Col- <br />orado. <br />10. The Gunnison River below Delta, <br />Colorado, .to its mouth. <br />11. The Colorado )livcr in Glen Canyon, <br />immediately below Glen Canyon Dam. <br />12. The Colorado Hiver in Marble and <br />Grand Canyons, Arizona. <br />Most of thc study area consists of can- <br />yons; however, major differences exist in <br />fish habitat. Areas 5, 6, and the upper <br /> <br />;.'f:~ <br />