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<br />
<br />HOLDEN AND STALNAKER-COLORADO RIVER FISHES
<br />
<br />219
<br />
<br />Many of the streams in the upper basin
<br />flow free, but two major reservoirs were
<br />built in the 1960's: Baming Gorge Reser-
<br />voir located on the upper Green River and
<br />Lake Powell just upriver from Marble and
<br />Grand Canyons (Fig. 1).
<br />Evermann and Rutter (1895) summarized
<br />collections from the Colorado River made
<br />prior to 1894. Taba, Murphy, and Frost
<br />(1965) made collections in the Colorado
<br />River for several miles below Moab, Utah,
<br />and Smith (1959) and McDonald and Dot-
<br />son (1960) conducted preimpoundment sur-
<br />veys in Glen Canyon. A fish eradication
<br />program in September 1962 (Binns et al.
<br />1964) effectively eliminated most native
<br />species above and for many kilometers
<br />below Flaming Gorge Dam before closure
<br />in November 1962. Vanicek, Kramer, and
<br />Franklin (1970) documented the distribution
<br />of native fishes below this dam following
<br />closure, and summarized preimpoundment
<br />surveys. They found no reproduction of na-
<br />tive fishes in the cold tail waters of the
<br />reservoir for 104 kilometers downstream.
<br />At that point, the Yampa River joins the
<br />Green and ameliorates the coldness of the
<br />Green somewhat. Areas noi previously
<br />studied include the lower 400 km of the
<br />Green River, the Yampa River, and the
<br />Colorado River above Moab, Utah.
<br />Minckley and Deacon (1968) reported
<br />two squawfish taken in the Grand Canyon
<br />in the mid 1960's. No subsequent accounts
<br />of squawfish from this canyon have been
<br />published. Minckley (1973) reported the
<br />humpback sucker as nearing extinction
<br />helow Lake Mohave.
<br />Presently, Colorado squawfish and
<br />humpback chub are considered "En-
<br />dangered" by the U.S. Department of the
<br />Interior (1973). Humpback sucker and
<br />bony tail chub are also rare (Miller 1972).
<br />The present study was undertaken to de-
<br />termine the status of fishes in the middle
<br />and upper Colorado River basins, with
<br />special emphasis on endemic, large-river
<br />forms.
<br />The objectives were: (1) to determine
<br />distribution and relative abundance of
<br />fishes in the Colorado River basin above
<br />
<br />Lake Mead, and (2) to identify areas im-
<br />portant for reproduction of endangered
<br />forms.
<br />
<br />STUDY AREA
<br />
<br />That part of the Colorado River basin
<br />(Fig. 1) considered in this study includes
<br />the Green River system from Craig, Col-
<br />orado; the Yampa River to the junction of
<br />the Green and Colorado Rivers; and the
<br />Colorado River from Hifle, Colorado to
<br />Lake Mead, excluding Lake Powell. Sev-
<br />eral larger tributaries also included were
<br />the Gunnison River below Delta, Colorado,
<br />and the Dolores, Price, Whitc, and Little
<br />Snake Rivers.
<br />The main system was divided into 12
<br />sampling areas:
<br />1. The Upper Yampa River from Craig,
<br />Colorado, to Juniper Springs, Colorado,
<br />approximately 80 km.
<br />2. The Lower Yampa Hiver in Yampa
<br />Canyon, Dinosaur National Monument, ap-
<br />proximately 72 km.
<br />3. The Green River from the mouth of
<br />the Yampa River to the south boundary of
<br />Dinosaur National Monument, about 32
<br />km.
<br />4. The Green River from Ouray to
<br />Green River, Utah, including Desolalion
<br />and Grey Canyons, about 192 km.
<br />5. The Green River from Mineral Bot-
<br />tom to its mouth, most of which is in
<br />Canyonlands National Park, about 104 km.
<br />6. The Colorado River for 16 km up-
<br />stream from mouth of the Green River.
<br />7. The Colorado River upstream from
<br />Moab, Utah, approximately 32 km.
<br />8. The Colorado River from Grand Junc-
<br />tion to Fruita, Colorado, about 16 km.
<br />9. The Colorado River near Rifle, Col-
<br />orado.
<br />10. The Gunnison River below Delta,
<br />Colorado, .to its mouth.
<br />11. The Colorado )livcr in Glen Canyon,
<br />immediately below Glen Canyon Dam.
<br />12. The Colorado Hiver in Marble and
<br />Grand Canyons, Arizona.
<br />Most of thc study area consists of can-
<br />yons; however, major differences exist in
<br />fish habitat. Areas 5, 6, and the upper
<br />
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