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<br />. <br /> <br />relationships that are biologically meaningful. Predictive outputs are not <br />supported by field observations. <br /> <br />In the Yampa River, where fish are successfully spawning every year and where <br />adults reside year-round, it can be assumed that the existing natural <br />conditions are conducive to success of endangered fishes. Model outputs for <br />the species and life stages examined herein do not predict flow/habitat optima <br />that even remotely approximate natural flow conditions, and do not fit within <br />the framework of a normal hydrograph. <br /> <br />The SI curve is the presumed biological link between fish and the available <br />and usable stream habitat. It is the state-of-the-art of human attempts to <br />simplify and quantify nature. Of the three variables sampled, water velocity <br />is probably the most influential factor for spawning and adult habitat <br />selection. The model output is also the most sensitive to changes in the <br />shape of the velocity curve. Depth is also important, however, after <br />exceeding some minimum, depth may not greatly influence habitat selection by <br />target species. loosely-compacted gravel is probably important for spawning, <br />although the other uses of gravel as substrate for invertebrates. and/or as <br />cover, need to be more clearly defined and understood. Cover requirements of <br />adult Colorado squawfish and humpback chub may dominate their selection of <br />microhabitat. Biological needs may quickly shift from protected low-velocity <br />habitats for resting areas, to higher velocity, deeper water which may afford <br />optimal feeding positions. The preservation of a balance of these habitats <br />does not appear practicable with present technology without substantive inputs <br />of empirical biological information. <br /> <br />Stream flow modeling using PHABSIM is more of an art form than a pure science. <br />Predictive model output (simulations) can be no better than the hydraulic <br />data. species criteria, and the ability of the model user to calibrate. select <br />the appropriate software application. and interpret results. However, the <br />need to quantify habitat for fish and wildlife use is increasingly important <br />as greater demands are put on an already limited water supply. PHABSIM can be <br />used as a management tool to assess some physical changes in habitat as a <br />result of altered streamflow. However, the link between the physical model <br />and the biological needs of endangered fish species needs significant <br />improvement before it can be used as a basis for determining instream flow <br />requirements for the Yampa River. Further, we believe that future field data <br />collections should focus on critical areas and/or limitin9 habitats rather <br />than on "representative reaches II so that collection of more detailed. high- <br />quality, site specific data is feasible. <br /> <br />Finally. we believe that a holistic approach to endangered fish management is <br />essential. and the species (or life stages) selected for study should reflect <br />the environmental constraints on the population as a whole. Formulating <br />stream flow recommendations for adults is meaningless if channel stability, <br />spawning, larval transport exotic competition and/or other factors are <br />limiting to the population of target fishes. An estimate of WUA as a function <br />of discharge is not an end in itself. Unless this index can be related to the <br />success of endangered fishes in some quantifiable way. it should not be used <br />as the basis for a State water right filing for protection of endangered <br />fishes. <br /> <br />18 <br />