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<br />riparian species like plains cottonwood must keep <br />pace with a decl ining alluvial water table. In <br />southern Alberta, Mahoney and Rood (1991) suggest <br />that survival of riparian poplar seedlings is most <br />likely in years where drawdown rates do not exceed 4 <br />em/day. Results of the above studies indicate that <br />cottonwood establishment and early growth are likely <br />to be successful provided drawdown rates are not <br />greatly in excess of 4 em/day and base flow <br />conditlons do not cause alluvial groundwater to <br />retreat below approximately 1 m. <br /> <br />Longer-term Survival <br /> <br />Bradley and smith (1986) found that recruitment or <br />establlshment of indivldual cottonwood stands on <br />river point-bars occurred approximately once in every <br />5 years. Although peak flows sufficient to support <br />establishment occurred once in levery 2 years, <br />coincidence of peak flows with; seed dispersal <br />occurred less freq!-lently. Lon<;{er"7te~ sur;vival of <br />cottonwood stands 1S also assoc1ated w1th lmportant <br />fluvial geomporhic processes. Along the Little <br />Missouri River in North Dakota, Everit (1968) found <br />that age and areal distribution of.~ottonwood stands <br />were closely associated with patterns of channel <br />movement and flood plain building processes. When a <br />river sandbar or point-bar attaLned a height of <br />approximately 1.5 m above low water, cottonwood <br />seedlings became established on . the bar surface. <br />Typically, seedlin~s in such locations are subject to <br />"chronic flood-tralning" (Lindsey et al. 1961) as a <br />result of sediment deposition during subsequent high <br />flows. Flood-trained saplings are flattened in a <br />downstream direction by flood waters and buried by <br />freshly deposited sediments. New stems arise from <br />the flood-trained stem but are again flattened and <br />buried by floods, giving them a stair-step <br />appearance. With continued deposition and movement <br />of the channel away from the point-bar, a stem <br />eventually escapes the zone of flood-training as <br />fewer floods inundate the aggrading surface. As a <br />stem continues to increase in girth it tends to <br />resist the force of a moderate flood, and is likely <br />to be uprooted and swept away only by a very large <br />flood. <br /> <br />THE H'iDROGRAPH <br /> <br />Germination and establishment of riparian vegetation <br />occur in response to the character and pattern of <br />streamflow w1thin a given year or growing season. <br />Consequently, an annual hydrograph (as compared to a <br />mean annual hydrograph) best represents and defines <br />those critical hydrologic elements that meet the <br />ecological requirements of riparian plants. A mean <br />annual hydrograph may obscure the unique behavior of <br />a stream or river within any given year. The <br />specific timing, magnitude, and intensity of an <br />individual peak flow, for example, are muted or lost <br />when averaged over an extended period of time. <br />Figure 1, therefore, depicts an annual hydrograph of <br /> <br />240 <br />