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<br />INTRODUCTION <br /> <br />Riparian zones or bottomlands are water courses and <br />associated areas of alluvial sediments. In arid and <br />semi-arid regions of North America, riparian <br />woodlands bordering streams and rivers are <br />distinctive features representing topographic <br />positions that are consistently moist enough to <br />support the growth of trees in a .landscape that is <br />otherwise too dry. Riparian zones represent important <br />transitional areas between upland and aquatic <br />ecosystems (Gregory et ale 19~1) and provide <br />essential habitat for a wide diversity of vertebrate <br />species (Brinson et ale 1981, Knopf 1985). <br /> <br />Western riparian ecosystems, particularly at lower <br />elevations, are often dominated by~species of po~lar <br />and willow. These early successional speC1es, <br />including plains cottonwood (Populus deltoides subsp. <br />monilifera (Aiton) Eckenwalder) 'are adapted to <br />relatively frequent, intense disturbances produced by <br />the energy of flowing water in riparian environments <br />(Osterkamp and Costa 1987). streamflow in most large <br />western rivers and streams is now influenced br <br />management activities including agricultura <br />diversion and damming for power generation and flood <br />control. Such activities have; caused dramatic <br />changes in the diversity, structure~ and organization <br />of riparian vegetation throughout western North <br />America. Changes in the magnitu~e and timing of <br />flows have led to marked declineslin the extent of <br />riparian vegetation in some river systems (Johnson et <br />ale 1976, Fenner et ale 1984, Rood.and Mahoney 1990). <br />However, in situations where natural flow <br />fluctuations historically prevented establishment of <br />vegetation, the moderating e.ffects of water <br />development have led to at least tem~orary increases <br />in the extent of riparian vegetat10n (Nadler and <br />Schumm 1981, Williams and Wolman 1984). <br /> <br />Ha,nagement of riparian resources within a larger <br />framework of water management will require clear <br />articulation of flow characteristics relevant to <br />riparian vegetation. In this paper we discuss stream <br />hydrograph characteristics critical to conditions <br />required for establishment, growth, and survival of <br />native riparian vegetation. <br /> <br />ECOLOGICAL REQUIREMENTS <br /> <br />The capacity of moving water to erode, transport, and <br />deposit alluvial materials is central to the <br />structuring and maintenance of western riparian <br />ecosystems (Brinson et ale 1981). Early successional <br />species, like many riparian willows and poplars, <br />possess a number of ecological characteristics that <br />make them particularly well-suited to physical <br />conditions in riparian zones. We discuss the <br />ecological requirements for germination, <br />establishment, growth, and long-term survival of <br />riparian vegetation, using plains cottonwood as an <br />example. <br /> <br />238 <br />