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<br />130 Swimming Against the Current <br /> <br />In June 1979 the USFWS and USBR signed <br />a memorandum of understanding (MOU) <br />whereby the USBR agreed to fund a com- <br />prehensive investigation of the endangered <br />Colorado River fishes in the upper basin. The <br />USFWS agreed to obtain the information essen- <br />tial to providing biological opinions on im- <br />pacts of existing and proposed water-develop- <br />ment projects. The primary objective of the <br />USFWS'S effort, named the Colorado River <br />Fish Project (CRFP), was to acquire informa- <br />tion needed to recover endangered fishes <br />while allowing the USBR to operate existing <br />projects as well as to plan and construct pro- <br />posed projects, <br />Initially, the CRFP extended over 965 km of <br />the Colorado and Green rivers and their tribu- <br />taries in Colorado and Utah. Studies focused <br />on identification of spawning requirements, <br />habitat requirements of young and adults, mi- <br />gratory behavior, interspecific competition, <br />predation and food habits, effects of tempera- <br />ture, salinity, and chemicals, development of <br />culture techniques, disease and parasite diag- <br />nostics, and taxonomy (w. H. Miller et at. <br />1982d), Fieldwork from 1979 to 1981 em- <br />phasized sampling the upper Colorado River <br />from Lake Powell to Rifle, Colorado (Valdez <br />et at. 1982b), and the Green River from its <br />confluence with the Colorado upstream to <br />Split Mountain Gorge (Tyus et at. 1982b), to <br />determine distribution, relative abundance, <br />movements, and habitats of various life stages. <br />The 1979 MOU was amended in 1981 to <br />include an investigation of humpback chubs <br />in the Little Colorado River, Arizona (Kaed- <br />ing and Zimmerman 1983), and to expand <br />field studies in the upper basin to include the <br />Dolores and Gunnison rivers, Colorado-Utah <br />(Valdez et at. 1982a). Additional funds pro- <br />vided by the U.S. Bureau of Land Manage- <br />ment to include White River, Colorado-Utah <br />(w. H. Miller et at. 1982b), and funds from <br />Congress and the U.S. National Park Service <br />(USNPS) supported field research on the Yampa <br /> <br />and Green rivers in Dinosaur NM (W H. Mil- <br />ler et at. 1982c). Funding provided through <br />the Windy Gap process was used for a three- <br />year habitat use and radiotelemetry study of <br />Colorado squawfish and humpback chubs on <br />the upper Colorado River, and to investigate <br />use of "nonflow alternatives" (habitat devel- <br />opment, fish passage, and stocking) as means <br />of maintaining and ensuring recovery of the <br />fishes of concern (Archer et at. 1985). <br />Laboratory research for Colorado squaw- <br />fish outlined in the MOU included swimming <br />stamina, bioassays of potentially toxic trace <br />element tolerances, and determinatiofl<<)f pref- <br />erences for temperature and total dissolved <br />solids. In addition, various contracted studies <br />examined physicochemical habitat conditions <br />in the Green and Colorado rivers, culture of <br />rare fishes, diseases, movements of Colorado <br />squawfish in the inlet to Lake Powell, and <br />stomach contents of fishes (W. H. Miller et at. <br />1982a), <br />The accumulated knowledge of the endan- <br />gered Colorado River fishes was summarized <br />in a 1981 symposium that emphasized srudies <br />conducted after 1975 (W. H. Miller et at. <br />1982C). The CRFP continued its work after <br />1982, emphasizing filling gaps in knowledge <br />of ecological requirements of the four rare <br />fishes; for example, delineating necessary <br />stream flows, describing movements (espe- <br />cially those associated with reproduction), <br />and identifying factors limiting recruitment <br />(Archer et at. 1984; Kaeding et al. 1986; Tyus <br />et at. 1987). <br /> <br />Other Studies <br /> <br />Studies of the ecology of rare fishes were also <br />conducted by biologists from the Colorado <br />Division of Wildlife (CODOW) and the Utah <br />Division of Wildlife Resources (UTDWR) from <br />the early 1980s to the present (e.g., Haynes <br />and Muth 1982; Radant 1982, 1986; Wick et <br />al. 1985). This work, funded through section <br />