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<br /> <br />'.r <br />'. <br />Ii <br /> <br />~ <br /> <br />Introduction <br /> <br />239 <br /> <br />or riparian vegetation, and the ability of soil to absorb and retain water <br />(Baker et aI., 1988). <br />Periodic flooding of aquatic ecosystems is a natural phenomenon to <br />which native flora and fauna are adapted, although organisms can be ad- <br />versely affected by catastrophic events. Ecological functioning of many <br />wetlands depends on periodic flooding or recharge from groundwater. <br />The riparian zone around lakes can become flooded when water inflow <br />is greater than outflow, but this is generally of short duration without <br />serious effects to plants and animals. Tide fluctuations and flows from <br />streams and rivers cause dynamic natural changes in salinity, water tem- <br />perature, nutrient transfer, and sedimentation in estuaries. Catastrophic <br />flooding of seacoasts occurs from hurricanes and tidal waves (tsunamis) <br />caused by earthquakes or volcanic eruptions. Natural catastrophic flood- <br />ing in large rivers can occur from formation and failure of lava dams <br />deposited by volcanoes, as in the Grand Canyon of the Colorado River <br />in southwestern United States (Hamblin, 1990), or failure of ice dams <br />that occurred during the Pleistocene in the northwestern United States <br />(Baker. 1981). The power associated with such flooding can dramati- <br />cally change landscapes (Baker and Costa, 1987). For example, the fail- <br />ure of glacial Lake Missoula in the northwestern United States released <br />as much as 2000km3 of water with peak discharges of 15-17 million m3 <br />S-l that were 30 thousand times greater than that produced by the largest <br />river on Earth, the Amazon River in South America (Baker and Bunker, <br />1985). <br />Flooding in small headwater streams is generally minor compared with <br />flooding in larger streams and rivers because drainage areas of the former <br />are smaller. Small streams are often found in forested landscapes where <br />soils can absorb and retain rainfall and meltwater. Flash flooding occurs <br />in small streams of arid regions when a sudden torrent of turbulent water <br />and sediment from summer thunderstorms in mountains causes erosion of <br />canyons, gulches, and arroyos. These streams are generally ephemeral so <br />aquatic ecosystems do not become permanently established and damage <br />is usually physical. <br />Generally, flooding of the piedmont reaches of large rivers is more com- <br />mon than in other aquatic systems because of the large volume of water <br />collected from an entire drainage basin. This water may cover a large area <br />of floodplain. Floods in large rivers can result in significant effects on <br />landscapes and aquatic systems. <br />Resiliency of most aquatic ecosystems damaged by flooding is often very <br />rapid. For example. catastrophic flooding of streams may result in the loss <br />ofa year-class of trout (Seegrist and Gard, 1972; Hanson and Waters. 1974; <br />Hoopes; 1975). However, Hanson and Waters (1974) documented the rapid <br />recovery in the standing crop and production rate of brook trout (Salvelinus <br />fontinalis) in a flood-damaged stream. In California. survival of O-age brook <br />trout fa fall spawner) was lowin years when fall floods occurred and high <br />in years after spring floods (Seegrist and Gard, 1972). Seegrist and Gard <br />reported that this pattern was reversed for spring-spawning rainbow trout, <br />Oncorhynchus mykiss. in the same stream. <br />This chapter discusses (1) the ecological importance of flooding to <br />aquatic ecosystems with an emphasis on large river systems, (2) effects <br />of human perturbations on aquatic ecosystems, (3) a case review of water <br /> <br />