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<br />254 <br /> <br />The Southwestern Naturalist <br /> <br />vol. 36, no. 2 <br /> <br />could be due to phenotypic variation, the rela- <br />tively smaller sizes and young age of chub col- <br />lected from these areas, or hybridization with <br />other Gila species. <br />The Little Snake River exemplifies the vari- <br />ation in discharge under which native fishes <br />evolved in the Colorado River System. The can- <br />yon in the lower 12 km of the Little Snake River <br />represents important local habitat for native fish- <br />es and should be protected. Natural flow regimes <br />should be maintained in the Little Snake River <br />to preserve this potentially important tributary <br />habitat for native fishes. Useful information on <br />distribution, taxonomy, and behavior of endan- <br />gered fishes may be obtained from further study <br />of the Little Snake River and other tributaries in <br />the Upper Colorado River System. <br />Funds to conduct radiotelemetry surveys were <br />provided by the United States Bureau of Recla- <br />mation, Salt Lake City, Utah. Funding for field <br />collection of fish and discharge measurements was <br />provided by the Colorado Division of Wildlife <br />through Study No. SE-3-12, Endangered Fishes <br />Investigations. H. Tyus and K. Paulin of the <br />United States Fish and Wildlife Service, Vernal, <br />Utah, contributed equipment and field assistance. <br />E. Wilson and R. Teller of the United States <br />Geological Survey, Meeker, Colorado, provided <br />discharge data for the Little Snake River. H. <br />Copeland and D. Wick assisted in field collec- <br />tions. R. Behnke, K. Bestgen, C. Carlson, and R. <br />Muth provided helpful reviews during prepara- <br />tion of the manuscript. <br /> <br />LITERATURE CITED <br /> <br />HAYNES, C. M. 1980. Endangered humpback chub <br />range extension documented. Endangered Species <br />Tech. Bull., 5(10):35. <br />HOLDEN, P. B. 1973. Distribution, abundance, and <br />life history of the fishes of the Upper Colorado River <br />Basin. Unpubl. Ph.D. dissert., Utah State Univ., <br />Logan, 59 pp. <br />HOLDEN, P. B., AND E. J. WICK. 1982. Life history <br />and prospects for recovery of Colorado squawfish. <br />Pp. 98-108, in Fishes of the Upper Colorado River <br />system: present and future (W. H. Miller, H. M. <br />Tyus, and C. A. Carlson, eds.). Amer. Fish. Soc., <br />Western Div., Bethesda, Maryland, 131 pp. <br />McKEOWN, B. A. 1984. Fish migration. Timber <br />Pres!J, Be'averton, Oregon, 224 pp. <br />SEETHALER, K. 1978. Life history and ecology of the <br />Colorado squawfish PtychocheiLus Lucius in the Up- <br />per Colorado River Basin. Unpub1. M.S. thesis, <br />Utah State Univ., Logan, 156 pp. <br />UGLAND, R. C., B. J. COCHRAN, R. G. KRETSCHMAN, <br />E. A. WILSON, AND J. D. BENNETI. 1989. Water <br />resourCeS data, Colorado, water year 1988, V 01. 2. <br />Colorado River Basin. U.S. Geol. Surv. Water-data <br />Rept., CO-88-2:1-370. <br />VALDEZ, R. A., AND G. H. CLEMMER. 1982. Life <br />history and prospects for recovery of the humpback <br />and bony tail chub. Pp. 109-119, in Fishes of the <br />Upper Colorado River system: present and future <br />(W. H. Miller, H. M. Tyus, and C. A. Carlson, <br />eds.). Amer. Fish. Soc., Western Div., Bethesda, <br />Maryland, 131 pp. <br /> <br />-..., <br /> <br />, . <br /> <br />.. <br /> <br />,. <br />