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<br />DISCUSSION. Rhinichthys osculus moapae is characterized by <br />adaptations to s'wift water. Large, expansive, falcate dorsal and anal <br />fins and expansive, deeply-forked caudals are typical. The caudal <br />peduncle is relatively long and slender. ;\iIany of these swift-water <br />adaptations are illustrated by comparing a spring form of Rhinichthys <br />05culus (for example, the population of R. o. robustus inhabiting the <br />Big Shipley Spring, Diamond Valley, Eureka County, Nevada) to <br />R. o. moapae (Table 1). Rhinichthys osculus moapae exhibits larger <br />size, more slender head, body, and caudal peduncle, greater anal to <br />caudal length, and greater dorsal and caudal fln lengths. Complete re- <br />sults of morphometric and meristic analyses are reported in Tables <br />2. and 3. <br />In addition to the information provided in the tables, the presence <br />or absence of barbels and premaxillary frenum was recorded. Of 32- <br />fish examined for these characters, 90.6% had 2. barbels and 93.8% <br />had a premaxillary frenum, which often is well developed. No fish <br />vvere found with only 1 barbel. All fish with barbels also possessed a <br />premaxillary frenum and only 1 fish without barbels possessed a pre- <br />maxillary frenum. Pelvic fins were absent in 4.1 % of 74 fish examined. <br /> <br />RELATIONSHIPS AND DIAGNOSIS. Hubbs (1943) interpreted <br />a subspecies as any genetic form which shows reasonable geographical <br />or ecological consistency, and which can usually be distinguished on <br />its totality of characters. By these criteria, R. o. moapae, with its nu- <br /> <br />TABLE 1 <br /> <br />l'dorphometric comparison of Rhinichthys osculus moapae with R. o. robustus from <br />Big Shipley Spring, Diamond Valley, Nevada. lWean (in parentheses) <br />and range are given. <br /> <br />R. o. rolJustus* <br /> <br />R. o. moapae <br /> <br />Standard length (mm) <br />Head deptht <br />Predorsallength <br />Anal to caudal length <br />Greatest body depth <br />Caudal peduncle depth <br />Depressed dorsal fin length <br />Caudal fin length <br /> <br />31-37 (34) <br />185-218(198) <br />558-623(591) <br />303-354(330) <br />247-283(263) <br />127-154(136) <br />208-243(224) <br />246-292(259) <br /> <br />42-77 (53) <br />129-166(146) <br />481-575(510) <br />323-483(394) <br />183-253(214) <br />91-126(109) <br />223-298(258) <br />229-305 (271) <br /> <br />- DilLa from Hubbs ct aJ. (1974). <br />";- :\I(~asurements in thousandths of standa.rd length. <br /> <br />514 <br />