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<br />1036 <br /> <br />pulsed DC unit in April and May; however, a few <br />fish (N = 7) were taken later in the year in trammel <br />nets. <br />We anesthetized the fish with tricaine (MS-222, <br />200 mgIL), inserted beeswax-coated transmitters <br />into the body cavity via lateral incision, and closed <br />the wound with 2.0-mm nylon sutures of triple- <br />tied, individual surgical knots (Tyus and McAda <br />1984; Tyus 1988b). Four types of transmitters, <br />which ranged in weight from 11 to 23 g in air, <br />were used. All transmitters emitted unique signals <br />(frequency and pulse rate) in the 40.66~0. 700- <br />MHz frequency range. Theoretical transmitter life <br />(Tyus 1988a) ranged from 150 to 550 d. Radio <br />receivers were Smith-Root RF-40 and SR-40 <br />types. Radio-tracking was conducted from boat <br />and airplane by use of omnidirectional whip and <br />bidirectional loop antennas. Eleven fish were <br />tracked for more than I year (five fish were tracked <br />455-542 d; transmitters were reimplanted in six <br />other fish). This permitted a comparison of be- <br />havior in different years. <br />Radio-tracking detected fish that we and other <br />investigators had radio-tagged. Of 153 radio- <br />tracked fish (including 11 fish tracked for more <br />than 1 year), 119 were radio-tagged by me or an <br />assistant, and 34 were radio-tagged by others: 4 <br />by the Utah Division of Wildlife (S. R. Cranney, <br />personal communication); 6 by the Colorado Di- <br />vision of Wildlife (P. C. Martinez, personal com- <br />munication); 12 by Colorado State University (E. <br />J. Wick, personal communication); and 12 by Bioi <br />West, Incorporated (R. A. Valdez, personal com- <br />munication). Capture locations of radio-tagged <br />Colorado squawfish included the Yampa (N = 32), <br />Green (N = 101), White (N = 19), and Duchesne <br />rivers (N = 1). <br />Fish were tracked in the lower 552 km of the <br />Green River, lower 192 km of the Yampa River, <br />lower 200 km of the White River, and lower few <br />kilometers of the Duchesne River and other <br />smaller tributaries (Figure 1). The location of each <br />radio-tagged fish was fixed by triangulation (Bovee <br />1986; Tyus 1988a). Locations were recorded in <br />river kilometers (RK) upstream from the mouth <br />of each river, and general habitat type and sub- <br />strate (direct observation and feel) were recorded <br />at the apparent signal source (Priede 1980). Mi- <br />crohabitat data, including water depth (direct <br />measurement with a rod) and water velocity <br />(Marsh-McBimey current meter at 0.6 of the depth <br />below the water surface) were recorded during the <br />spawning season for fish that remained in one lo- <br />cation for 30 min or longer. Ifa located fish could <br /> <br />....... <br /> <br />~~ <br /> <br />TYUS <br /> <br /> <br />FIGURE I.-Green and Yampa rivers, where move- <br />ments and migrations of Colorado squawfish were stud- <br />ied, 1980-1988. <br /> <br />not be disturbed by wading or probing after the <br />30-min period, triangulation was repeated. If, af- <br />ter a second attempt, fish movement could not be <br />obtained, the record was not used. Data collected <br />from airplane flights were not used to determine <br />habitat. <br />Radio-tagged fish were classified according to <br />their movements during an approximate spawn- <br />ing season (June 15-August 1) as follows: mi- <br />grants-fish moving to one of two known spawn- <br />ing areas during the spawning period; suspected <br />migrants-individuals moving to an area consid- <br />ered suitable for reproduction during the spawn- <br />ing period (similar substrate and habitat to known <br />spawning areas); unknown-fish moving 30 km <br />or more to areas in which no spawning area was <br />suspected; sedentary-fish exhibiting only local <br />movements (<30 km) in which no migratory pat- <br />tern was evident; lost-fish whose signal was lost <br />after a month or less of tracking, or fish with which <br />no contact was made for a month or more during <br />the spawning season; and late implants-fish ra- <br />dio-tagged during or after the spawning season. <br />Tagging and recapture. -Colorado squawfish <br />longer than 350 mm total length (TL) captured in <br />the Green River basin by USFWS in 1979-1988 <br /> <br />r, . <br /> <br />"" <br /> <br />'1 <br />.. <br />