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<br />for endangered fishes can proceed using <br />combinations of empirical studies, analyt- <br />ical models, computer simulations, and <br /> <br />subjective assessments to assist the deci- <br />sion-making process (Maguire 1986; Soule <br />1987). <br /> <br />CONCLUSIONS <br /> <br />Instream flow requirements for endan- <br />gered fishes should be determined with <br />respect to their recovery, not survival. This <br />is particularly true in the Colorado River <br />where habitat change has been severe and <br />widespread, and is continuing. Survival is <br />too often synonymous with "status quo," <br />and this may ultimately lead to extinction <br />in declining populations. In the past, <br />stream flows for fisheries resources have <br />been provided on a minimum flow basis, <br />and the utility of this approach is ques- <br />tionable (Stalnaker 1990). <br />Although much progress has been made <br />in developing analytical methods for as- <br />sessing streamflows, the relationship be- <br />tween physical parameters and fish produc- <br />tion is complex and not well understood. <br />New approaches for assessing streamflow <br />requirements of fishes are needed if en- <br />dangered fish recovery is to become a re- <br />ality. Rather than providing endangered <br />fish with the same habitats in which they <br />are declining, alternatives might include <br />studies that would test fish community re- <br />sponse to various flow scenarios. These <br />could include appropriately designed lab- <br />oratory studies, reclamation of stream sec- <br />tions, or construction of new stream sec- <br />tions in which such things as flows and <br />species compositions could be manipulat- <br />ed to test specific hypotheses concerning <br />fish response to flows. The goal of these <br />studies should be to determine if flows are <br />limiting the recovery of different life stages <br />of the endangered fishes and to evaluate <br />the role of "non flow" impacts. Because ex- <br />perimental approaches are complex and <br /> <br />costly, they will not become a reality un- <br />less managers are better informed about <br />shortcomings and potential dangers of us- <br />ing some instream flow methods. <br />Although most of the interest in deter- <br />mining and protecting Colorado River <br />fishes has been associated with the need <br />to protect federally listed endangered spe- <br />cies, the "fundamentally insular" (Molles <br />1980) nature of the fauna suggests that oth- <br />er species may also become endangered or <br />extinct in the near future. As each species <br />disappears, it is anticipated that recovery <br />of the remaining forms will become in- <br />creasingly more difficult as perturbations <br />in the native ecosystem increase. <br />Successful recovery of the Colorado Riv- <br />er fishes to a self-sustaining status will re- <br />quire the efforts of many individuals with <br />varied interests concerning multiple uses <br />of water. Because some of these interests <br />will differ, progress will be made only by <br />concentrating on the consequences of wa- <br />ter management decisions. Such decisions <br />must be based on a recovery flow philos- <br />ophy grounded in sound ecological prin- <br />ciples and solid biological information. <br /> <br />Acknowledgments <br /> <br />I thank the Bureau of Reclamation, Fish and <br />Wildlife Service, National Park Service, and other <br />agencies and universities that have supported re- <br />search and management activities on endangered <br />Colorado River fishes. Additionally, I thank C. A. <br />Karp, J. Hamill, and three anonymous reviewers <br />for their comments on earlier drafts of the manu- <br />scri pt. <br /> <br />REFERENCES <br /> <br />Baltz, D. M., B. Vondracek, 1. R. Brown, and P. B. Moyle. 1987. Influence of temperature on <br />microhabitat choice by fishes in a California stream. Transactions of the American Fisheries <br />Society 116(1):12-20. <br />Baxter, G. T., and J. R. Simon. 1970. Wyoming fishes. Cheyenne: Wyoming Game and Fish <br />Department (Bulletin 4). <br />Beecher, H. A. 1990. Standards for instream flows. Rivers 1(2):97-109. <br /> <br />I H. M. Tyus <br /> <br />33 I rt <br />