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REVIEWS in FISHERIES SCIENCE Editor Robert R. Sticlzney University of Washington Seattle, Washington Editorial Advisory Board Robert Anderson Chesapeake Biological Laboratory Solomons. MD Claude E. Boyd Department of Fisheries and Allied Aquaculture Auburn University Auburn, AL Kevin Fitzsimmons Environmental Kesearch Laboratory University of Arizona Tucson, AZ John Kraeuter Shellfish Laboratory Rutgers University Por[ Norris, NJ Takeshi Murai Bio-function Division National Research Institute of Fisheries Science Yokohama, Japan Richard A. Neal National Marine Fisheries Service La Jolla, CA Eric Hallerman Department of Fisheries and Wildlife Virginia Polytechnic Institute Blacksburg, VA Ronald Hardy Nc» •thwest Fisheries Center Seattle, WA Anne Henderson-Arzapalo Fish Culture Research Laboratory Kerneysville, WV Harold L. Schramm Mississippi Cooperative Fish and Wildlife Research Unit Mississippi State, MS Sandra Shumway Marine Resources Laboratory West Boothhay Harbor, ME Patrick Sorgeloos State University of Ghent Ghent, Belgium Renews in F'islaeries Science is published by CRC, Press, Inc., 2000 Corporate Blvd., N.W., Boca Raton, FL 3343L USA. For 4 issues, U.S. rates are $79.95 to individuals; $195.00 to institutions; add $7.95 per issue foreign shipping and handling fees to all orders not shipped to a United States or Canada zip code. For immediate service and charge card sales, call our toll free number. 1-800-272-7737 Monday through Friday, 7:30 a.m.-6:00 p.m. EST (Continental U.S. and Canada only). To order by FAX: 1-800-374-3401. Or send written orders to: CRC PRESS, INC., P.O. Box 750, Pearl River, NY 10965- 0750. This journal contains information obtained from authentic and highly regarded sources. Reprinted material is quoted with permission, and sources are indicated. A wide variety of references aze listed. Reasonable effors have been made to publish reliable data and information, but the editor and the publisher cannot assume responsibility for the validity of all materials or for the consequences of their use. All rights reserved. Awhorization to photocopy items for intemal or personal use, or the personal or intemal use of specific clients, may be granted by CRC Press, Inc., provided that $.5(l per page photcxopied is paid directly m Copyright Clearance Center, 27 Congress Street, Salem, MA, 01970 USA. The fee code for users of the Transactional Reporting Service is ISSN 1064-1262/93 $0.00 + $.50. The fee is subject [o change without notice. For organizations that have been granted a photocopy license by the CCC, a separate system of payment has been arranged. CRC Press, Inc.'s consent does not extend to copying for general distribution, for promotion, for creating new works, or for resale. Specific permission must be obtained from CRC Press for such copying. ISSN I(I(v1-1262 No claim to original U.S. Government works ©1993 by CRC Press, Inc. Boca Raton Ann Arbor London Tokyo Printed on acid-free paper