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<br /> <br />· 'f <br />1 <br />, <br />\ <br /> <br />394 <br /> <br />J, A, STANFORD ET AL. <br /> <br />FLOODPLAIN <br /> <br />HILLSLOPE I <br /> <br />TERRACE <br /> <br />I <br /> <br />Pioneer <br /> <br /> <br />r~" F"j' <br /> <br />Upland Forest <br /> <br />Climax <br />Riparian Forest <br /> <br /> <br />Channel <br /> <br /> T <br /> ..... <br /> . <br /> ..... <br /> 0 <br /> 3 <br />,. 1 <br />5-1 0 km I <br /> <br />Figure I, Major landscape features of a montane floodplain river, showing the three primary spatial dimensions (lateral, longitudinal or <br />altitudinal, and vertical) that are dynamically molded through time (the fourth dimension) by fluvial processes, Biota may reside in all <br />three spatial dimensions: riparos (streamside or riparian), benthos (channel), hyporheos (river-influenced groundwater) and phreatos <br />(deep groundwater). The hatched area is the varial zone or the area of the channel that is periodically dewatered as a consequence of <br />the average amplitude of the discharge regime. Major channel features include a run (A), riffle (B) and pool (C); Sd refers to sites of <br />sediment deposition and Se refers to a major site of bank erosion. The heavy solid line is the thalweg and broken lines conceptualize <br />circulation of water between benthic, hyporheic and phreatic habitats (after Stanford, 1996; see also Stanford and Ward, 1992) <br /> <br />GENERAL PRINCIPLES OF RIVER ECOLOGY <br /> <br />Conservation and management strategies for large rivers must have a solid conceptual basis or they will <br />likely fail to sustain biodiversity and bioproduction, Contemporary river ecology is guided by a number <br />of intertwined concepts or principles derived from empirical studies. No two rivers are exactly alike and <br />no single theory encompasses the myriad of biophysical interactions and responses to natural and human <br />disturbances that make each river unique, However, fundamental principles do apply; many conservation <br />and restoration efforts become myopic, costly and too often fail because plans and actions overlook ecolo- <br />gical fundamentals. <br />Unregulated rivers exist as geohydraulic continua from continental divides to the oceans. They are net- <br />works of surface and groundwater flow paths that drain catchment landscapes (Gibert et at., 1990), The <br />energy of flowing water constantly reconfigures the physical form of these interconnected flow pathways, <br />primarily by the process of cut and fill alluviation (Leopold et at" 1964) although dissolution can dominate <br />in limestone massifs (Mangin, 1994) and a few other situations, Inorganic and organic materials are eroded <br />upstream and deposited downstream primarily in relation to: (a) long- and short-term flow dynamics; (b) the <br />resistivity of geological formations to erosion and dissolution; (c) instream retention structures (e.g. eddies, <br />wood debris); and, (d) the geometry of the catchment. <br />Channel morphologies are determined by the legacy of flooding. Big floods fill channels with inorganic <br />and organic materials eroded laterally and vertically at upstream locations, thereby producing a continuum <br />of instream structures (pools, runs, riffles, gravel bars, avulsion channels, islands, debris jams) and lateral <br />floodplain terraces in many sizes and shapes, Local morphologies resulting from infrequent, very large floods <br />may persist in the same general form (quasi-equilibrium) for long time periods until the next big flood, even <br />