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<br /> <br />lydro- <br />a can- <br />iOSS of <br />r even <br />pulses <br />ic and <br />low or <br />cannot <br />1t clear <br />rmour- <br />he can- <br />:r time, <br />1 et ai., <br />vegeta- <br />ply and <br /> <br />vers the <br />n; habi- <br />channel <br />gure 5). <br />st rivers <br />riverine, <br />always <br />j Leidy, <br />tal core <br />gulation <br />ams and <br />that use <br />sing the <br />3). <br />lutrients, <br />ain inun- <br />opportu- <br />)86; Li et <br />ew native <br />pervasl ve <br />ive inver- <br />~aches (Li <br />ich opens <br />e riparian <br />lOd exotic <br />chi, 1994; <br />[) general, <br />lS the fact <br /> <br />')wnstream <br />) explicitly <br />, described <br />'ivers, The <br />~ontinuum <br />ic state) of <br /> <br />RESTORATION OF REGULATED RIVERS <br /> <br />403 <br /> <br />the reservoir, the mechanics of water release (surface, bottom or depth selective), the mode of dam <br />operations and the influence of tributaries entering downstream from the dam. If the tributaries are large <br />and unregulated, they may substantially mediate the reset (Stanford and Hauer, 1992). In any case, <br />conditions at some point downstream from the dam will closely approximate conditions elsewhere in the <br />continuum. Thus, upstream or downstream shifts in biophysical conditions mediated by dams manifest as <br />predictable discontinuities in the river continuum. For example, biophysical conditions at some <br />predictable point downstream from a large bottom release (hypolimnial) dam in the montane transition <br />of a temperate latitude river will be very similar to pristine conditions far upstream, because of the cool, <br />clear water released from the reservior. In rivers that are free flowing for long distances downstream from <br />large dams in the montane reaches, the position of the rhithron-potomon transition can be predicted <br />from the operational mode of the dams relative to the influence of tributaries, <br />The predictions of the SDC along the longitudinal dimension have been largely substantiated (Stanford et <br />ai., 1988; Ward and Voelz, 1988; Hauer et ai" 1989; Stanford and Ward, 1989; Ward and Stanford, 1990, <br />1991; Munn and Brusven, 1991; Sabater et ai" in press), although recent incorporation of responses of large <br />floodplains (Ward and Stanford, 1995b) require additional resolution, The main point is that the ecological <br />consequences of specific regulation schemes are largely predictable, and environmental degradation asso- <br />ciated with regulation can be ameliorated. We recognize that uncertainties derive from interactions with pol- <br />lution and the introduction of exotic biota. However, pollution can be curtailed or eliminated, and non- <br />native biota are likely to be substantially less successful as invaders when dams are operated in ways that <br />maximize resets of environmental heterogeneity, <br /> <br />RESTORA nON PROTOCOL <br /> <br />The era of dam building may be over in much of the world because high efficiency and affordable dam sites <br />are already developed, Loss of biodiversity and bioproduction, especially riverine and anadromous fisheries <br />(Frissell, 1993; WeIcomme 1995). underscores the need for restoration of regulated rivers and enormously <br />expensive reconstructions are underway or are being planned (Dahm et ai" 1995; Gore and Shields, <br />1995), Even removal of large dams on large rivers is included in some restoration plans because the costs <br />of damage to fisheries and other attributes of riverine integrity in some instances far exceed the commercial <br />value of the dams. Removal of large dams is obviously problematic in a variety of ways, such as the mobi- <br />lization of large volumes of fine sediments stored in the reservoir basin, and methods for evaluating removal <br />strategies have been proposed (Shuman, 1995), A variety of approaches exist for restoring small streams with <br />substantial emphasis on engineered structures such as weirs, off-channel ponds, rock gardens (Gore and <br />Shields, 1995) and many other artificial habitat structures (Hunter, 1991). Commercial operations advertise <br />engineering expertise for bulldozing damaged streams back to pre-regulation channel configurations and <br />stories of restored fisheries and improved water quality abound in the popular literature, although scientific, <br />long-term evaluations of such schemes are much less available (Sear, 1994), Structures placed in stream are <br />often washed out, fail to restore biodiversity or produce unanticipated negative responses, such as increased <br />bank erosion or accelerated deposition of fine sediments (Frissell and Nawa, 1992) and increased water tem- <br />peratures (c. Frissell, unpublished data) associated with weirs and rock gardens, Such problems largely <br />derive from lack of attention to the conceptual foundations of river ecology and the first principles of the <br />ecology of regulated streams, <br /> <br />Formalize the probiem at catchmen{ scale <br /> <br />Restoration of large, regulated rivers begins with recognition of the river continuum and evaluation of <br />the loss of ecosystem capacity to sustain biodiversity and bioproduction, Biological (e,g, past and present <br />distribution of native biota) and physical (e.g, channel configuration) indices of ecosystem resilience are <br />needed (Frissell et al., 1993); measures of biological integrity as defined by Angermeier and Karr (1994) <br />may be more useful than biodiversity per se because of the difficulty of accurately determining the distri- <br />bution and abundance of benthos, fish and other river organisms. Habitat requirements for all life history <br />stages and generation times (turnover rates) of native, keystone species (i,e" top carnivores and other <br /> <br />