<br />.
<br />
<br />the month of September for the Colorado River near the
<br />staleline in C0)nrado.
<br />
<br />Habitat Time Series
<br />
<br />From time series and duration analyses, the water man-
<br />agement community can determine the likelihood of given
<br />flow events occurring under various proposed regulation
<br />schemes. It therefore behooves the fishery manager to trans-
<br />late fishery habitat events into similar kinds of analyses. To
<br />do this for each river segment. one must first assemble flow-
<br />habitat functions such as those shown in Fig. 3 and 7, which
<br />represent the output from an instream water temperature
<br />macrohabitat model and the microhabitat model PHABSIM,
<br />respectively. With the hydrologic time series and the flow
<br />habitat functions, the dynamics of the physical habitat can
<br />be described for a specific river reach over time, while
<br />simultaneously illustrating various water management
<br />effects.
<br />Ageneralized equation for the habitat versus streamflow
<br />function,ean be written as follows:
<br />(4) HA, = /;<Q,)L
<br />where HA, is the quantity of usable physical habitat. Q is
<br />the streamflow. and fi(Q) is the functional relation illus-
<br />trated by Fig. 7. The subscript i refers to the specific fish
<br />species or life stage under investigation and L is the length
<br />of stream determined from a macro model (Fig. 3) to be
<br />represented by fi(Q) at time t.
<br />From a time series of flows for a specific river segment
<br />(Q,), the amount of usable physical habitat at time (t) can
<br />be calculated. The result of a unit length transformation of
<br />the fi(Q) function from Fig. 7 is illustrated in Fig. 8 for
<br />adult rainbow trout. The hydrologic time series for this river
<br />is also plotted in Fig. 8 for comparison. These time traces
<br />illustrate several important concepts that appear to be true
<br />for many fish species in many different types of streams:
<br />(1) the physical habitat time series is less variable than the
<br />streamflow time series; (2) high flow events may be as
<br />detrimental to fish as low flows; and (3) during some time
<br />periods, a moderate reduction in streamflow can result in
<br />a large reduction in physical habitat (e.g.. compare August
<br />1973 with August 1972) and other time periods when the
<br />converse is true (compare June 1971 with December 1973).
<br />Average monthly flow data were used to generate the hab-
<br />itat time series in Fig. 8. If daily flows were used and the
<br />
<br /> 9 45
<br />::- - PHYSICAL HABITAT
<br />,
<br />E --- STREAMFLOW
<br />.. en
<br />~ ....
<br />~ 6 30 :Il
<br /> m
<br />< I , >
<br />~ I \ :::
<br />iii ~ I , , , I
<br /> , I I , 'Tl
<br />< ,~. : , , , , I roo
<br />:I: I , , I , 0
<br /> ,,, I I I '" , ::E
<br />-' : \1 , , ,I I
<br />< , " I " , 15 3'
<br />U I I , , I { ,
<br /> I ,
<br />0, ' I I I I , !"
<br /> I 1,\ : I \ , I
<br />> I I , ..
<br />:I: ~' V I I I I
<br /> I
<br />Q. , I \ -
<br /> \ \
<br /> 'I I \
<br /> 0 0
<br /> 1971 1972 1973 1974
<br />
<br />WATER YEAR
<br />
<br />FIG, 7. Mean monthly streamflow and physical habitat for adult
<br />rainbow trout during 1971 through 1974 in the North Fork Sno-
<br />qualmie River, Washington.
<br />
<br />22
<br />
<br />~J
<br />
<br />
<br />e,;
<br />E
<br />-
<br />
<br /><
<br />w
<br />[[
<br />e( 4.5
<br />l-
<br />e(
<br />t:
<br />m
<br />e(
<br />J:
<br />g 1.5
<br />o
<br />~
<br />
<br />6
<br />
<br />- - - ~>- -
<br /> - .-
<br />I- -
<br /> - r--
<br /> -- - --
<br /> -- - c-
<br /> -
<br />
<br />3
<br />
<br />o
<br />
<br />1965
<br />
<br />1970
<br />
<br />1975
<br />
<br />1980
<br />
<br />WATER YEAR
<br />
<br />FIG, 8. Annual limiting microhabitat for adult rainbow trout in the
<br />North Fork Snoqualmie River, Washington.
<br />
<br />habitat results averaged to arrive at the average monthly
<br />habitat values, the monthly values would not be the same.
<br />The habitat values for one water year, based on daily and
<br />monthly data, are compared in Table 2. The differences are
<br />not large for the North Fork Snoqualmie River; however.
<br />they still illustrate how important it is to use the same trans-
<br />formation procedure when comparing water management
<br />options such as those for pre- and post-project conditions.
<br />The choice between monthly or daily streamflow hydrologi-
<br />cal series depends on the objective of the analysis, available
<br />data, and funds. For example, daily values could be used
<br />for a gaged site where the water resources system is being
<br />simulated by using daily flows. In contrast, monthly stream-
<br />flows would be used for a location with few existing stream-
<br />flow measurements and where the streamflow record must
<br />be synthesized by regression with nearby sites.
<br />Chronological time sequences are important for fishery
<br />studies because habitat analyses for all life stages are not
<br />applicable for every month of the year. For instance, rain-
<br />bow trout spawning may occur from April to June. Spawn-
<br />
<br />TABLE 2. Comparison of the results of different microhabitat gen-
<br />eration procedures for adult brown trout (Salmo trulla) in the
<br />North Fork Snoqualmie River, Washington, October 1972 to Sep-
<br />tember 1973.
<br />
<br />Month
<br />
<br />Oct.
<br />Nov.
<br />Dec.
<br />Jan.
<br />Feb.
<br />Mar.
<br />Apr.
<br />May
<br />June
<br />July
<br />Aug.
<br />Sept.
<br />
<br />Daily
<br />streamflows
<br />
<br />Monthly
<br />streamflows
<br />
<br />(Square metres per linear metre of stream)
<br />
<br />4.85
<br />5.86
<br />5.52
<br />6.41
<br />5.80
<br />6.62
<br />6,74
<br />6.53
<br />6.53
<br />4.79
<br />2.26
<br />3.48
<br />
<br />5.34
<br />6.89
<br />5.52
<br />6.34
<br />6.16
<br />6.98
<br />7.08
<br />6.50
<br />6.59
<br />5.19
<br />2.26
<br />4.79
<br />
|