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<br />Beckman, IWd; Si!.:ler and \Iilll'r, 19.13; :\Iilll'r and Low(', 196i).
<br />Ecolypl', a Il'rIll propo.';l'd h." TlI n',.;son (I ()2 2) ior plan l..;. ;IS "thl.'
<br />prodllct arisin:.: as a reslllt oi gl'llot."pical respOIlSl' of an eco-
<br />specil's to a particlIlar hahitat," is ,!.(l'llel;i11." I'qlli,'all'lll to :\lilln's
<br />LIS\' of lilt' krm "l'cological slIhspl'cies." The "I'cOSIll'cil's." also
<br />(oilll'd hy TlIrrssl)ll. is I'qlli,':dl'lll III tIll' "'LiIlIl:IL':11l specil's from
<br />'In ecological poim oi vie\\' I :\Iayr. )9(J3:3.111." .\layr t'/ 11/. (19.1,);
<br />313) ddille ('co,,;pl'cil's (ll1odiiil'd from TlIrl'sson) ,I.' "'agrollp of
<br />pOJllllations so rl'latl'd Ih;1l tIlt'." an' ahll' to I'\ch:lngl' gl'nl'S frl'l'ly
<br />withollt loss of fl'rtility or ,'igor ill oifspring'" alld l'l'olypl' as ";1
<br />dl'scripti\'(. tl'rlll applil'd to pl:lI11 racl'S of ,'arying dl'grl'l's of dis-
<br />tiIlCIIH'SS which 0"'1' llll.ir mosl COlhpicuou,,; ch:lractl'rislirs to llll'
<br />sl'!("("ti\'(, \'fil'Ct..; of Illcal l'll\,irollllll'llh." Earlil'r, IllIlllh (I (},.\.))
<br />,.;tall-d thaI "ill illtTl'a..;in:.: Illlmlll'rs. sllhsJll'cil',.; aI',' IJl'ill,g ,;llIJ\rll
<br />10 hI' l'colo,gical or lllicrogl'o~raJlhical racl's"' (S('I' also IllIhh..;, I ')(1 I).
<br />:\nolhl'r term th,lI ma." Ill' pertilll'nt is "l'cotypic "arialioll." as
<br />ddilll'd hy :\layr (1l)(13:-t15l, "the iormalion within each geo-
<br />!!raphic area oi IJIllll,'r'''I': loc;dized pop"l.rli"!,,, tklt sJll'cialtz,' J/l
<br />ecologically diiil'l"l'llt sullan'a,,;. alld are not olJlitl'ratl'd Il~' gelle
<br />('\chall~1' with adjacent. diiferl'ntl.\' spl'cialized pIlPll1:rlillllS." In
<br />appliclli'lll Ili slIch l"IlIlCl'pls tll thl' fisllt's 11I'I'SI'IlII.\' lIlld!'r COIl-
<br />~idl'ralilln (1'\r1l1dill~ (;. l'I,'~,"IS). till' l'co,.;pecil''; wllllld be Gill/
<br />I'o/I//sll/. with Ihn'!, I'ClllYPl''; Ill' I'CIlSlIlbl'l'cil':'. G. 1', I'o/J/lsl,/, (;, 1'.
<br />gralll/llli. all" (;, 1', illlallln/ill. Thl' "\'aryin~ d!.grl'('s Ili distinct-
<br />Ill'S,';" \\'Illlld Ill' I hl'i I' l.\tern:t! /11llrphlllll;':y, in a gradl'd snil's from
<br />(J', 1'. illl,nlln/ill, thrllll~h (;. 1'. ~I'I/lrl/II/i tll (;. I'oll//slll, as dl'..;nilJl'd
<br />Ilei\II'\',
<br />Ilal,it:Lls irlllll \rhirh I'hlllJ_, h:I\'I' 1Jl'l'n cllllt-Cll'" I,y Ill(' :Inrl IIlh..'r
<br />per.-IInnl'l irlllll .\ri/lln:l Stall' l'lli\"('r,i,~' an' gt.nnall.\' inclllhi,;-
<br />ll'nl wilh :In "\'colYpl'" illll'rp1'l'l:llillll. Fill' ('\:l111jl!e spl'Cilllens IIf
<br />G. r,';I'I/IrI/Jlli ha\'l' I'l'l'n 1:lkt'n in ,';111:111. \\.arlll, rlt'..;nl.;lrl'anlS likl'
<br />.\r;I\':lill:1 t'rn'/; (111<':111 ~r:lilil'nl 7.() 111 /;/11). \\.hich CllIlSi._I..; prin-
<br />cipally IIi riffll''; alld il'w pllllls (ILlrhn alld \linl'kll'Y. !l)(,(J) ,:Ild
<br />in Fllssil ('rcd; which is l::r~I' and swiil wilh :1 Clllhl:lIll-lI'lllp('ralllrl'
<br />'prin~ at its SlllllTl' and a drill' Ilf :t!rllllsl .'iCO III ill ilS IS km
<br />descellt tll 1111' \'erdl' Ri\'l'r. Thi.; slIIJSPI'cil'S abll j,; rd:\tively
<br />llniillrm In Ihl' IIppl>r (;iLr River 1J:I..;in. frll/11 Ihl' Iargl' /11ainstrl'alll
<br />10 Ihl' high mlll/11lain (2._'i00 III 3,OU(1 /11) Iriblltaril's.
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