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<br />l)b
<br />
<br />TilE \\':\S~IY'\~ JO\'R~.\1. OF III1H.llt;\", \",,1.. .\~, ~". I, 1',-',
<br />
<br />including G. r. gra/I/I/l/i, aho\"l' falls of small tributaries. ~Ion'm, l"
<br />of downstream populations. presllmahly (;. r. robns/lI by this tiL
<br />also was possible anrl they il1\'adl.d upstrt':lm anrl largely displ:i,
<br />the former isol:lll'. (;. r. grall/l/l/i.
<br />:\n oh,.jous "hematin' to the intnpr('\atil;ns gin'n so far is "
<br />cmdary contact and hyhridization (~[ayr, Il)().\: :;0:' I hl'l\\"el'11
<br />(;ila Ri\'er endelllic (G. il///"rlll/'l/io), and an uppn Ila..;in fl)rlll I
<br />rolll/s/Il). This cannot he cOlllph-tL'ly rejected. If tnll', hO\\"(',.,:
<br />the h\'hridization III II st predate clllling of the lake Ill'ds (ll1i,:
<br />(':11'1\' j'll'iston'IH') since {;. r. ~r(/I/(//l/i no,,' persists in hanging tril,:
<br />tari~'s (~'i:.. allll"l' Ilarril'r falls) of the \'t.'rdl' systl'lll (i.l'., /-"41' il
<br />Creek) and ,hll\\'" rl'll1a rka hll' hI Ill1ogenl.i t y in 1ll4Jrphology a nd I",
<br />ristics 0\'1'1' a din'rse spl'ctrulll of halJitah. and in n'l:ition 10 til\'-
<br />I'rl'sl'nt intl'rgradation hl'lwl'l'n tml suhspl'cil's ((;. r. rolll/s/lI :\!
<br />G. r. il/l(,/"/l/l'llill) also may ,gl'nerally Ill' rejl'clt-rI on the sallle ba,,'
<br />~ll cllrrent ZOlll'S of sympatry l)('twI'en G. rolli/sill and G. il/la/II/'rl/ll
<br />are known hilt SOllll' olrll'r records (I'.~.. (;ilbnt :lIld Scofield. ISI)S I.
<br />indicate local sYlllpatry in comjlh'" hahitats. The interllll'diatl'
<br />for III , G. grall/l/l/i, may show diif'.'rl'ntiation frolll place to plan',
<br />a possihll' indication of isolation aile.\"(, dallls of tl1l' different ri\"(."
<br />rlurin.g "lio-l'lt-istoCl'nl' ill1pOIlIHln1t'nt. Thesl' ,'ariations sel'lll incl,.
<br />pl'ndl'nt of thl' pro\illlily of (;. illla/l/o/i". \"ariation \\.ithin (,
<br />illll'/"I/I/'dill is highly l4Jcalizl'd (i./'.. .\14Jnkl'y ~pring"), and in tli,.
<br />GIS(' of (;. r. robn.ilo, S('l'IllS atlrihut:t1,II' tll intergr:"btion froll1 (;
<br />r. grtl!J"/l/i popllbtion.;, .01' pll\'notypic rl'S[lIln.;l' to local ('n\'irtll'
<br />Illental conditions.
<br />TIll' proposed nrigin of G, illlo'/l/oli" fr4J1ll thl' SOllth is "u:
<br />stantiated I,y ils appal'l'n\ rl'l:ltion.;hip" to chub,; oi Ih' .\ll'xi\:l\\
<br />l'latl'all. Rdat('ll ,pecil''; oi thl' adjacent. ,lllltlll'rIl and l'a,ll'I!!
<br />ba,;ins include (;, dila/'I/ia :'.Iilll'r irlllll thl' In.;in IIf the Rill dl' i!
<br />Concepcion. :\rizona and :-;onora, and tIll' Rio :-;onora. ~onora: (,
<br />lIi~rt".i,.,'m (( ;ir:trll) from the c'lllll'll'\ (;uZlllan ba"in, ~('W :'.Il'xi,.
<br />awl ('hihu:lhua: and (;. pal/tlllfll (( '0)11') irom thl' Rio (;r:lIldl' aI:'
<br />I'I'COS hasins, ~l'\\' ~ll'"ico and TI''':IS, ,\11 arl' Iargl'-scall'd, thi.;.
<br />hodil'd. darkly pignll'ntl'd ii,;lll's with rclatin'ly loll' scale and iin-r:
<br />counts (tallll' ,; l. quilt, unlike G. rO/.I/"la and it" rl'lati"I'';, :11
<br />\"ery similar to (;. il/lt"J"/l/t'llia, :\11 oi these, phh (;. /lJ"(l/lli (Eigl'l
<br />mann and Eigl'nmann) oi coaslal streams of soullll'rn ('aliforni:i,
<br />
<br />-."c,
<br />
<br />C\"I'RI~ID FISIIE~-RI:'\~E
<br />
<br />tJj
<br />
<br />were con..;idl'rl'd hy l'yeno (196() toha\"(' originalt'd frolll tilt'
<br />"ancestral forlll that .~a\'e rise to G. r. il//a/J/nlill in the Gila Rin'r."
<br />If population,; IIf (;. ill I alii n/ill an' mon' doseh' related to t hosl'
<br />oi till' south. as proposed hl're,thl'n they are m:)SI likl'h' derin'd
<br />frolll captures in,'oh'in,g north-ilowing trihutaric,; of the (;'ila Ri\'l'r.
<br />or perhaps l'arlin a,; part of an anCil'nl iauna of the ori,ginal ..;outh-
<br />II'l'st tn'lHling stn'alll ,;y';Il'Ill.';. 1 1 II\\"('\'er , shifling of Jand Illas,;('.;
<br />along coa,;t:d area,; IIi II'l',;II'm ~ll'\ico and Baja ('aliillrnia cannot
<br />he rllh-d out Cllllll'll'tely as a n1\':1I1S oi dispersal. Connt'ction,; IIi
<br />the IIppl'r (;ila Rin'r SI'.gnH'nt to the (;uzll1an ha,;in 01 northern
<br />Chihuahua (('olllt,y and /);l\.id,;on. 19(,3: \\"1.1 IeI'. in h.olllll,,',-ki t'I
<br />al.. 1l)(I';: .\ll'ltlln. 1")(1';' i" nllt :iul'pllrtl'd Ill' 1'\'idl'nCl' frolll the
<br />irhthYlIi:lllna, (;ila lIi~rt'sl'OIS IIi lhl' (;uzman i,asin i,; quite "imil:lr
<br />to (;, il/II'/"II//'tlia, difit'l"in~ nllhtly in ha\'ing l) dllrsal fin-rav,; ralhl'r
<br />than S. Yd, no SPl'ril'."i and il'W ,gl'nera are in COllllllon bl't-\\l'l'n lilt'
<br />(;ila and (;UZIll:ln systl'nh, Ihl' latter having principall\' a Rill
<br />Grande a,;s('ll1ll\;lgl' 01 nati\"(' iisl1l's, whereas thl' (;ila Ri;'l'r IJ:lsin
<br />is occupied hy a di';linctive. highly l'ndelllic faulla (~Iilll'l", Il)SS).
<br />One aquatic organislll. a unionid clam. .JI/OdOIlIIl t"lzliillfllit'l/sis Ll'a.
<br />knoll'n from a numlll'r oi western drainage,; a,; f~)s,;il or living
<br />material (T:lylllr, 19().l). is pre';l'nl in both the (;ila anrl (;uzm:lll
<br />ha,;ins. Its alJSl'nCl' frolll ;l(ljau'nl.,;olllherJl S\',;ll'nh mal. h()\\l'\'l'r
<br />reflect a lack IIi colll'cting (J), \\'. Taylor, fll'l";on:d Clllllll;,;nicllion,:
<br />The Rio Yaqui "-,"skill and olhl'r minor coastal drailla~~I's oi
<br />SOllora, \/l'xico. sharI' a 1l11l11l>er of ichthyofaun:tl l'lellll'nt,; with
<br />Ihl' (;ila (\I('el,;. II)O.l: .\lilkr. 19,;Sl. These are .Igosia d. (11/"\'10-
<br />,~asla ((;iranl), l'II",'ilill/lsis 1It"l'id/'l/lalis (Baird ;Ind (;ir;'trllJ. 'and
<br />olhl'r pairs oi ,;pl'cie,; that are c1o';l'ly rl'lall'd: ('almlllllil/,i illsi~l/i.\
<br />l!:iird :11111 (;irard and C, InTl/artlilli (;irard: (;. ro/'l/sl,1 :lnl1 (;.
<br />lliil/aOlt' .\II'('k ( (;. mIJJHla:': :'.Iilll'r 11)':;,-; J, and anothl'r (;ila
<br />that Il1:1Y 1)(' 11\':11' (;. inlaillnlia ("G. nigrc,\'(t'lIs" of \Il'l'k J 90-1. in
<br />part: dors:ll and :111:11 iin-rays S, scalI''; in thl' Iatl'ral sl'ril''; 60 i':;).
<br />:\Iso, a nUIl1IH'r oi Rio (;r:lIllll' fi..;hl''; h:ln' Jll'll\'tr:lll'd thl' Rill Yaqui
<br />(c.g" .YOlfOpis, ('lIi11po.lltlJlJ/l. l'iJiJl"plzall's. Ie/all/rl/s), pr('~ul1\al>h'
<br />frol11 tIll' (;UZI1I:1l1 h:l..;in :Inri or frol1l thl' Rio l'oncho,;, Thl' alJsl'nc'(,
<br />of 111\'..;e in lhl' (:ila Rin'r, l'sp('ciall.\" some of thl' mOrl' ag.!.!n',;"iw,
<br />upland minno\\"s. indicatl''; an invasion of the Yaqui that postllatb
<br />an.\" Iargl' faunal ('"change I>l'I\\"('en that drainagl' and thl' 111\\"('1'
<br />Color:lllo b:lsi n.
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