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1166 Is cs,siu, 111'diolq;ir.Werulion <br />system) or current conditions to simulated results <br />based on models of future modification to a system. <br />The data we use in estimating all attribute values are <br />daily mean water conditions (e.g., levels, heads, flow <br />rates). The same computational strategies will work <br />with any regular-interval hydrologic data, such as <br />monthly means, but the sensitivity of the IRA method <br />for detecting hydrologic alteration is increasingly com- <br />promised with time intervals longer than a day. Detec- <br />tion of certain types of hydrologic impacts, such as the <br />rapid flow fluctuations associated with hydropower.gen- <br />eration at dams, may require data from even shorter in- <br />tervals (e.g., hourly). <br />Hydrologic Attributes <br />Hydrologic conditions can vary in four dimensions <br />within an ecosystem (three spatial dimensions and <br />time). If the spatial domain is restricted to a specific <br />point within a hydrologic system, however, (such as a <br />measurement point in a river, a lake, or an aquifer), the <br />hydrologic regime can be defined in terms of one tem- <br />poral and one spatial dimension: changes in water con- <br />ditions (e.g., levels, heads, rates) at a single location over <br />time. Such temporal changes in water conditions are <br />commonly portrayed as plots of water condition against <br />time, or hydrographs (Fig. 1). <br />. Our goal is to characterize the temporal variation of <br />hydrologic conditions using attributes that are biologi- <br />cally relevant yet sensitive to human influences such as <br />reservoir operations, ground water pumping, and agri- <br />cultural diversions. Many attributes of hydrologic re- <br />gimes can be used to characterize the "physical habitat <br />templates" (Southwood 1977, 1988; Poff & Ward 1990; <br />Townsend & Hildrew 1994) or "environmental filters" <br />(Keddy 1992) that shape the biotic composition of <br />aquatic, wetland, and riparian ecosystems. The IRA <br />method is based on 32 biologically relevant hydrologic <br />parameters divided into five major groups to statistically <br />characterize intra-annual hydrologic variation (Table 1). <br />These 32 parameters are based upon five fundamental <br />characteristics of hydrologic regimes: <br />(1) The magnitude of the water condition at any <br />given time is a measure of the availability or suit- <br />ability of habitat and defines such habitat attributes <br />as wetted area or habitat volume, or the position of <br />a water table relative to wetland or riparian plant <br />rooting zones. <br />(2) The tinning of occurrence of particular water con- <br />ditions can determine whether certain life-cycle re- <br />quirements are met or can influence the degree of <br />stress or mortality associated with extreme water <br />conditions such as floods or droughts. <br />(3) The frequency of occurrence of specific water <br />conditions such as droughts or floods may be tied <br />kichlc'rl sir. <br />1600 <br />1400 <br />1200 <br />1000 <br />?E <br />Soo <br />E <br />W <br />600 <br />400 <br />200 <br />0 <br />1600 <br />1400 <br />1200 <br />1000 <br />800 <br />E <br />600 <br />400 <br />200 <br />0 <br />Month <br />Figure 1. Two hydrographs for the Roanoke River at <br />Roanoke Rapids in North Carolina can be cbaracter- <br />ized by the five general features of a hydrologic re- <br />gime: magnitude, frequency, duration, tinning, and <br />rate of change. These regime features can be altered by <br />human influences sucli as dams, as illustrated by a <br />comparison of the.upper pre-danz 15ydrograph for <br />1942 with the lower post-danz hj?drograph for 1975 <br />(cins = cubic meters per second = 35.315 cubic feet <br />per second). <br />to reproduction or mortality events for various <br />species, thereby influencing population dynamics. <br />(4) The duration of time over which a specific water <br />condition exists may determine whether a particu- <br />lar life-cycle phase can be completed or the degree <br />Consrnation 6iolop <br />Volume Ill. No. -t. August 19% <br />J F M A M .1 J A S O N D <br />.1 F M A M .1 J A S b N D