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1726 D. B. OSMUNDSON ET AL. Ecological Applications <br />Vol. 12, No. 6 <br /> 250 8 100 <br /> a -Width 7 <br /> 200 4 R ©Depth E <br />6 0 80 <br /> <br />150 ? <br />/ `-' <br />5 <br />N <br />60 <br />3 ? <br />a V <br /> 100 3 C 40 <br /> != <br />2 U <br />U 50 U 20 <br /> 1 <br />0 <br />0 0 <br />10- <br />8 <br />°E 6 <br />t0 <br />4 <br />N <br />2 <br />2 <br />LU <br />16 <br />V 12 <br />0 <br />8 <br />4 <br />n <br />C <br />U <br />25 <br />E <br />N 20 <br />A <br />N 15 <br />U <br />10 <br />a <br />O 5 <br />0 <br />?N 0 <br />100 <br />80 <br />rn <br />C 60 <br />N <br />'Q <br />40 <br />E <br />1L 20 <br />n <br />lb <br />V 14 <br />O 12 <br />U <br />C)_ 10 <br />U) <br />N 8 <br />r-. 6 <br />O <br />4 <br />Q <br />N 2 <br />O <br />n <br />U. U6 <br />N 0.07 <br />E <br />0.06 <br />N 0.05 <br />E 0.04 <br />O <br />> 0.03 <br />v <br />'O 0.02 <br />0.01 <br />0 <br />1 2 3 4 5 6 7 8 91011 1 2 3 4 5 6 7 8 9 1011 <br />Stratum Stratum <br />Fic. 2. Longitudinal pattern of physical parameters by study stratum: (a) bankfull width and depth of channel; (b) <br />percentage of surface area of the dominant habitat types (run and riffle); (c) mean D50 particle size of riffle and run substrates; <br />(d) mean D50 particle size of riffle and run substrates excluding particles <2 mm; (e) fraction of riffle and run surface material <br />consisting of particles <2 mm; (f) percentage of embeddedness (aerial surface area composed of fines) of riffle and run <br />substrates; (g) depth of free space within riffle and run substrates; (h) volume of void space within riffle and run substrates. <br />Values plotted are means ± 1 SE.