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<br />688 <br /> <br />OSMUNDSON ET AL. <br /> <br /> <br />.,I1,/" <br /> <br />~L- <br />lC' <br /> <br />0-<- <br />{v "if <br />~ <br />~<<:- <br /><1 <br />'v <br /> <br />6 <br />(/>.:. <br />'tP <br />(j <br />",-0< <br />(} <br /> <br />UTAH <br /> <br />UPPER REACH <br /> <br />o <br />, <br /> <br />25 <br /> <br />50 75 <br /> <br />100 <br /> <br />KILOMETERS <br /> <br />COLORADO <br /> <br />FIGURE I.-Map of Ihe upper Colorado River study area. Transverse Jines indicate boundaries of upper and lower <br />study reaches (Westwater Canyon was not studied). Upstream-most lines mark sites of physical barriers to fish <br />movement; no such barriers exist at other Jine locations. <br /> <br />Problems inherent in scale-based aging and <br />growth estimation for long-lived species, such as <br />Colorado squawfish, prompted us to refine existing <br />growth estimates by using new mark-recapture <br />data. Age estimates so derived also enabled us to <br />assess rates of adult survival. Our primary objec- <br />tive was to refine estimates of Colorado squawfish <br />growth rate by minimizing use of scale analyses <br />and relying instead on recaptured PIT-tagged fish. <br />A secondary objective was to estimate adult sur- <br />vival rate with an approach modified from models <br />(e.g., Seber 1982) that use declining numbers of <br />increasingly older fish in the population. A third <br />objective was to estimate age of Colorado squaw- <br />fish of various sizes and provide an estimate of <br />potential longevity. In addition, we discuss the dis- <br />appearance of very large individuals reported in <br />historical accounts and mortality factors that may <br />at present reduce survival rates. <br /> <br />Methods <br /> <br />Study Area <br /> <br />The study area included the occupied range <br />(about 300 km of river) of the Colorado squawfish <br />in the Colorado River upstream of the Green River <br />confluence (Figure 1). Locations are in river ki- <br />lometers (rk) from the Green River confluence (rk <br />0.0), converted from river miles mapped by Belk- <br />nap and Belknap (1974). Further upstream move- <br />ment is blocked seasonally by a dam at rk 298.1 <br />and all year by another dam at rk 303.0. Upstream <br />access to the Gunnison River, a major tributary <br />entering at rk 275.1, is blocked all year by a dam <br />3.5 km upstream of the mouth. Most adult Colo- <br />rado squawfish live in the upper part of the study <br />area; earlier life stages are generally found down- <br />stream. The study area was divided into upper and <br />lower reaches based on distribution of life stages.. <br /> <br />" <br /> <br />. <br />