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' flow is determined from a data series <br />of discharges defined over a specific <br />time interval, and it has a frequency <br />of occurrence of 0.5 (a 50% prob- <br />ability). <br />*The duration is the period of time <br />associated with a specific flow condi- <br />tion. Duration can be defined relative <br />to a particular flow event (e.g., a flood- <br />plain may be inundated for a specific <br />number of days by a ten-year flood), <br />or it can be a defined as a composite <br />expressed over a specified time period <br />(e.g., the number of days in a year <br />when flow exceeds some value). <br />*The timing, or predictability, of <br />flows of defined magnitude refers to <br />the regularity with which they occur. <br />This regularity can be defined for- <br />mally or informally and with refer- <br />ence to different time scales (Poff <br />1996). For example, annual peak flows <br />may occur with low seasonal predict- <br />ability (Figure 2b) or with high sea- <br />sonal predictability (Figure 2c). <br />•The rate of change, or flashiness, <br />refers to how quickly flow changes <br />from one magnitude to another. At <br />the extremes, "flashy" streams have <br />rapid rates of change (Figure 2b), <br />whereas "stable" streams have slow <br />rates of change (Figure 2a). <br />Hydrologic processes and the flow <br />regime. All river flow derives ulti- <br />mately from precipitation, but in any <br />given time and place a river's flow is <br />derived from some combination of <br />surface water, soil water, and ground- <br />water. Climate, geology, topogra- <br />phy, soils, and vegetation help to <br />determine both the supply of water <br />and the pathways by which precipi- <br />tation reaches the channel. The wa- <br />ter movement pathways depicted in <br />Figure 3a illustrate why rivers in <br />different settings have different flow <br />regimes and why flow is variable in <br />virtually all rivers. Collectively, over- <br />land and shallow subsurface flow <br />pathways create hydrograph peaks, <br />which are the river's response to <br />storm events. By contrast, deeper <br />groundwater pathways are respon- <br />sible for baseflow, the form of deliv- <br />ery during periods of little rainfall. <br />Variability in intensity, timing, <br />and duration of precipitation (as rain <br />or as snow) and in the effects of <br />terrain, soil texture, and plant evapo- <br />transpiration on the hydrologic cycle <br />combine to create local and regional <br />Y• <br />?E <br />g' <br />P <br />C, <br />E <br />o" <br />Rte, <br />?4a "lee <br />v <br />b„ <br />_Y <br />?E <br />a <br />a? <br /> <br /> <br />r? <br />?x <br />Figure 2. Flow histories based on long-term, daily mean discharge records. These <br />histories show within- and among-year variation for (a) Augusta Creek, MI, (b) <br />Satilla River, GA, (c) upper Colorado River, CO, and (d) South Fork of the <br />McKenzie River, OR. Each water year begins on October 1 and ends on September <br />30. Adapted from Poff and Ward 1990. <br />flow patterns. For example, high <br />flows due to rainstorms may occur <br />over periods of hours (for permeable <br />soils) or even minutes (for imperme- <br />able soils), whereas snow will melt <br />over a period of days or weeks, which <br />slowly builds the peak snowmelt <br />flood. As one proceeds downstream <br />within a watershed, river flow reflects <br />the sum of flow generation and rout- <br />ing processes operating in multiple <br />small tributary watersheds. The travel <br />time of flow down the river system, <br />combined with nonsynchronous tribu- <br />tary inputs and larger downstream <br />channel and floodplain storage ca- <br />pacities, act to attenuate and to <br />dampen flow peaks. Consequently, <br />annual hydrographs in large streams <br />typically show peaks created by wide- <br />spread storms or snowmelt events <br />and broad seasonal influences that <br />affect many tributaries together <br />(Dunne and Leopold 1978). <br />The natural flow regime organizes <br />and defines river ecosystems. In riv- <br />ers, the physical structure of the en- <br />vironment and, thus, of the habitat, <br />is defined largely by physical pro- <br />cesses, especially the movement of <br />water and sediment within the chan- <br />nel and between the channel and flood- <br />plain. To understand the biodiversity, <br />production, and sustainability of <br />river ecosystems, it is necessary to <br />appreciate the central organizing role <br />played by a dynamically varying <br />physical environment. <br />The physical habitat of a river <br />includes sediment size and heteroge- <br />neity, channel and floodplain mor- <br />phology, and other geomorphic fea- <br />tures. These features form as the <br />available sediment, woody debris, <br />and other transportable materials are <br />moved and deposited by flow. Thus, <br />habitat conditions associated with <br />channels and floodplains vary among <br />December 1997 771