' flow is determined from a data series
<br />of discharges defined over a specific
<br />time interval, and it has a frequency
<br />of occurrence of 0.5 (a 50% prob-
<br />ability).
<br />*The duration is the period of time
<br />associated with a specific flow condi-
<br />tion. Duration can be defined relative
<br />to a particular flow event (e.g., a flood-
<br />plain may be inundated for a specific
<br />number of days by a ten-year flood),
<br />or it can be a defined as a composite
<br />expressed over a specified time period
<br />(e.g., the number of days in a year
<br />when flow exceeds some value).
<br />*The timing, or predictability, of
<br />flows of defined magnitude refers to
<br />the regularity with which they occur.
<br />This regularity can be defined for-
<br />mally or informally and with refer-
<br />ence to different time scales (Poff
<br />1996). For example, annual peak flows
<br />may occur with low seasonal predict-
<br />ability (Figure 2b) or with high sea-
<br />sonal predictability (Figure 2c).
<br />•The rate of change, or flashiness,
<br />refers to how quickly flow changes
<br />from one magnitude to another. At
<br />the extremes, "flashy" streams have
<br />rapid rates of change (Figure 2b),
<br />whereas "stable" streams have slow
<br />rates of change (Figure 2a).
<br />Hydrologic processes and the flow
<br />regime. All river flow derives ulti-
<br />mately from precipitation, but in any
<br />given time and place a river's flow is
<br />derived from some combination of
<br />surface water, soil water, and ground-
<br />water. Climate, geology, topogra-
<br />phy, soils, and vegetation help to
<br />determine both the supply of water
<br />and the pathways by which precipi-
<br />tation reaches the channel. The wa-
<br />ter movement pathways depicted in
<br />Figure 3a illustrate why rivers in
<br />different settings have different flow
<br />regimes and why flow is variable in
<br />virtually all rivers. Collectively, over-
<br />land and shallow subsurface flow
<br />pathways create hydrograph peaks,
<br />which are the river's response to
<br />storm events. By contrast, deeper
<br />groundwater pathways are respon-
<br />sible for baseflow, the form of deliv-
<br />ery during periods of little rainfall.
<br />Variability in intensity, timing,
<br />and duration of precipitation (as rain
<br />or as snow) and in the effects of
<br />terrain, soil texture, and plant evapo-
<br />transpiration on the hydrologic cycle
<br />combine to create local and regional
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<br />Figure 2. Flow histories based on long-term, daily mean discharge records. These
<br />histories show within- and among-year variation for (a) Augusta Creek, MI, (b)
<br />Satilla River, GA, (c) upper Colorado River, CO, and (d) South Fork of the
<br />McKenzie River, OR. Each water year begins on October 1 and ends on September
<br />30. Adapted from Poff and Ward 1990.
<br />flow patterns. For example, high
<br />flows due to rainstorms may occur
<br />over periods of hours (for permeable
<br />soils) or even minutes (for imperme-
<br />able soils), whereas snow will melt
<br />over a period of days or weeks, which
<br />slowly builds the peak snowmelt
<br />flood. As one proceeds downstream
<br />within a watershed, river flow reflects
<br />the sum of flow generation and rout-
<br />ing processes operating in multiple
<br />small tributary watersheds. The travel
<br />time of flow down the river system,
<br />combined with nonsynchronous tribu-
<br />tary inputs and larger downstream
<br />channel and floodplain storage ca-
<br />pacities, act to attenuate and to
<br />dampen flow peaks. Consequently,
<br />annual hydrographs in large streams
<br />typically show peaks created by wide-
<br />spread storms or snowmelt events
<br />and broad seasonal influences that
<br />affect many tributaries together
<br />(Dunne and Leopold 1978).
<br />The natural flow regime organizes
<br />and defines river ecosystems. In riv-
<br />ers, the physical structure of the en-
<br />vironment and, thus, of the habitat,
<br />is defined largely by physical pro-
<br />cesses, especially the movement of
<br />water and sediment within the chan-
<br />nel and between the channel and flood-
<br />plain. To understand the biodiversity,
<br />production, and sustainability of
<br />river ecosystems, it is necessary to
<br />appreciate the central organizing role
<br />played by a dynamically varying
<br />physical environment.
<br />The physical habitat of a river
<br />includes sediment size and heteroge-
<br />neity, channel and floodplain mor-
<br />phology, and other geomorphic fea-
<br />tures. These features form as the
<br />available sediment, woody debris,
<br />and other transportable materials are
<br />moved and deposited by flow. Thus,
<br />habitat conditions associated with
<br />channels and floodplains vary among
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