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DISPERSAL PATTERNS OF COLORADO SQUAWFISH <br />700 <br />650 <br />E 600- <br />E <br />550 1993-1994 <br />rn <br />C <br />500 <br />f>3 <br />0 450 <br />1991-1992 <br />400 <br />350 <br />1 2 3 4 5 6 7 <br />Stratum <br />FIGURE 2.-Mean length of Colorado squawfish by stra- <br />tum. Lengths of fish captured in 1991 and 1992 (heavy <br />line) and those captured in 1993 and 1994 (light line) were <br />pooled. Error bars represent 95% confidence intervals. <br />Results <br />Distribution <br />Mean length of Colorado squawfish captured in <br />this study progressively increased upstream (Fig- <br />ure 2). Because samples from strata 2, 3, and 5 <br />were small, lengths of fish captured were pooled <br />into 1991-1992 and 1993-1994 periods for statis- <br />tical tests. Analysis of variance (ANOVA) re- <br />vealed that for 1991-1992 data, mean lengths in <br />strata 3, 5, 6, and 7 differed significantly from the <br />mean in stratum 1; those in strata 5-7 differed from <br />the mean in stratum 2; mean lengths in strata 7 <br />and 3 also differed ((ANOVA, Tukey-Kramer <br />multiple comparison; P < 0.05). Regression in- <br />dicated significant relationships (Figure 3) be- <br />tween total length and rkm in 1991 (r2 = 0.59, P <br />< 0.0001) and 1992 (r2 = 0.60, P < 0.0001); <br />slopes between 1991 and 1992 were not signifi- <br />cantly different (P = 0.28). <br />For 1993-1994, differences in mean lengths <br />among strata were not as pronounced as in 1991- <br />1992. Significant differences were found only be- <br />tween stratum 1 and strata 3, 5, 6, and 7 (ANOVA, <br />Tukey-Kramer multiple comparison, P < 0.05). <br />Mean lengths in the two most downstream strata <br />were significantly greater during 1993-1994 than <br />during 1991-1992 (t-test, P < 0.0001). In the up- <br />per reach, mean lengths declined from 1991-1992 <br />to 1993-1994 (significantly only in stratum 7; <br />t-test, P = 0.012). Regression indicated significant <br />relationships between total length and rkm loca- <br /> <br />i uuv <br />800 <br />600 <br />400 <br />200 <br />0 <br />1000 <br />800 <br />E 6o0 <br />E 400 <br />200 <br />1) <br />C o <br />(1) 1000 <br />(ti 800 <br />0 <br />600 <br />400 <br />200- <br />0 <br />1000- <br />800- <br />600- <br />400- <br />200- <br />947 <br />1991 $ <br />$+++ <br />N=89 + <br />+ <br /> + <br /> <br /> <br /> r-squared = 0.59 P < 0.0001 <br /> <br />1992 <br />N=91 <br /> <br /> + <br />+ <br />+ <br />+ r-squared = 0.60 P < 0.0001 <br /> <br />1993 ++ + <br />N= 160 <br />+ <br />+ <br /> + <br />+ <br /> r-squared = 0.22 P < 0.0001 <br />1994 <br /> + <br />N = 127 <br />+ <br /> <br /> r-squared = 0.32 P < 0.0001 <br />0 30 60 90 120 150 180 210 240 270 300 <br />River kilometer <br />FIGURE 3.-Relationship between Colorado squawfish <br />length and kilometers upstream from the lower end of the <br />study area (river kilometer 0 at the Green River conflu- <br />ence) during 1991-1994. <br />tion in 1993 (r2 = 0.22, P < 0.0001) and 1994 (r2 <br />= 0.32, P < 0.0001). Slopes were not significantly <br />different between 1993 and 1994 (P = 0.13), but <br />both differed from slopes of relationships for 1991 <br />and 1992 (all P < 0.003). <br />Netting catch rates indicated that adult distri- <br />bution changed during 1991-1994. During 1991- <br />1992, very few adult Colorado squawfish were <br />caught in the lower reach. In 1991, Colorado <br />squawfish 500 mm and longer were captured only <br />in stratum 3 (one fish) and none in strata 1 or 2, <br />resulting in an overall mean CPUE in the lower <br />reach of 0.02 fish (?500 mm) per net (SE = 0.02; <br />N = 47 net sets). In contrast, 42 fish 500 mm or <br />longer were captured with nets in the upper reach, <br />for a mean CPUE of 0.30 (SE = 0.07; N = 139). <br />A similar pattern was evident in 1992: Colorado <br />squawfish 500 mm or longer were captured only <br />in stratum 3 (six fish) and none in strata 1 or 2.