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25 shows the river-wide averages for detritus densities in riffles and runs in the Colorado and <br />Gunnison rivers. No significant differences were found between the detritus levels in riffles <br />and runs in the Colorado River for the spring of 1994 and 1995. However, riffles were found <br />to be significantly greater in fall 1994 than in fall 1995. In both years, the amount of detritus <br />in riffles increased between the spring and fall periods while decreasing within run habitats. <br />The highest mean concentration of detritus occurred in riffles and the lowest concentrations <br />occurred within runs in fall 1994. <br />In the Gunnison River, detritus levels were highly variable (note the large 95% confidence <br />intervals in Figure 25) with mean values greater than the Colorado River for both runs and <br />riffles. However, these higher mean values were not significantly different from the mean <br />Colorado River levels. <br />Densities of secondary consumers were estimated by measuring the standing crop of <br />macroinvertebrates within the benthic substrate. These results are presented in Figure 26 for <br />the Colorado and Gunnison rivers. Riffles had significantly higher biomass densities <br />(expressed as a dry weight per area) when compared to run habitats. The highest densities <br />occurred in spring 1994 for both habitat types while the lowest densities occurred in spring <br />1995. The temporal pattern between the two habitat types was almost identical except during <br />fall 1994. Although both habitats experienced decreases relative to the spring 1994 densities, <br />the decrease was much greater in runs (1.9 gm/m2 to 0.43 gm/m2) versus riffles (4.1 gm1m2 <br />to 2.9 gm/m2). The biomass decrease in runs was significant (P<0.05) level while the <br />decrease in riffles was not significant. <br /> COLORADO RIVER-RIFFLES COLORADO RIVER-RUNS <br /> Mean Periphyton Chlorophyll-a Mean Periphyton Chlorophyll-a <br />100 100 <br /> <br />r <br /> <br />R 10 R 10 - <br />U V <br /> <br /> y <br />1 1 <br /> Spring 1994 Fall 1990 Spring 1995 Fall 1995 - Spring 1994 Fall 1994 Spring 1995 Fall 1995 <br /> PERIOD PERIOD <br /> GUNNISON RIVER-RIFFLES GUNNISON RIVER-RUNS <br /> Mean Periphyton Chlorophyll-a Mean Periphyton Chlorophyll-a <br />too 100 <br /> - <br /> <br />E E <br />W 10 - P 10 <br />O O <br /> <br />1 t <br /> Spring 1994 Fall 1994 Spring 1995 Fall 1995 - Spring 1994 Fall 1994 Spring 1995 Fail 1995 <br /> PERIOD PERIOD <br />Figure 24. The seasonal distribution of periphyton chlorophyll for riffles and runs in the Colorado River (above) and the <br />Gunnison River (below). <br />24