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in the Colorado River just prior to the spring 1995 sample period. <br />Biological Parameters-In order to define the trophic structure of the Colorado and Gunnison <br />rivers, primary and secondary producers were quantified in the same locations where physical <br />and chemical parameters were measured. <br />Primary producers were quantified by <br />determining the in-situ standing crop of <br />periphyton (mg Chl-a/m2) and the <br />concentration of benthic detritus <br />(gm/m2) associated with the substrate. <br />Periphyton biomass as expressed as <br />river-wide mean concentrations were <br />significantly different for each sample <br />period (Figure 12). Initial samples <br />collected in the Colorado River in the <br />spring of 1994 had river-wide average <br />concentrations of 7.2 mg/m2; this <br />increased to 9.8 mg/m2 by the fall of <br />that year. However, by spring 1995, the <br />periphyton densities were reduced to the <br />lowest levels (4.6 mg/m2) of the entire <br />study period. In the following sample <br />period (fall 1995) periphyton densities <br />had risen to their highest concentrations <br />(13.4 mg/m). This increase from the <br />spring to the fall of 1995 was the largest <br />increase found in the Colorado River <br />between sample periods. <br />Figure 12. The mean and 95% confidence interval of the <br />periphyton chlorophyll values calculated for the river-wide <br />substrate samples collected in 1994 and 1995 in the Colorado <br />(above) and Gunnison (below) rivers. <br />The Gunnison River also had significant differences between sample periods for periphyton <br />biomass (Figure 12) and had the same temporal pattern as the Colorado River, with the <br />spring of 1994 having the lowest densities and the fall of 1995 having the highest. In <br />addition, the fall 1994 and fall 1995 periphyton densities were significantly greater in the <br />Gunnison River than in the Colorado River. <br />The concentrations of coarse particulate organic material (detritus) in the substrates of the <br />Colorado River were not significantly different among sample periods (Figure 13) averaging <br />21-23 gm/m2. The Gunnison River, although having higher average detritus concentrations <br />for each respective sample date, was not significantly greater, with the exception of fall 1994. <br />15