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<br />1' <br />BACKGROUND <br />Floodplains have an influential role on river ecology (Bayley 1995, Ligon et al. 1995, <br />Townsend 1996). The annual hydrologic cycle of a floodplain consists of a periodic cycle of <br />inundation and recession, typically associated with seasonal weather patterns. River ecosystem <br />productivity has been shown to be limited by floodplain availability in some instances (Newbold <br />et al. 1983, Tiessen et al. 1994). In large river systems, significant available portions of nitrogen <br />and phosphorous are derived from floodplain habitats. They influence food web productivity <br />through the input of limiting nutrients from the bottom up in a hierarchical manner (Newbold et <br />al. 1983, Vought et al. 1994, Leonardson et al. 1994). Nutrients in solution, derived from <br />floodplain detritus and soils, are available for autochthonous production (photosynthesis) in the <br />growth of phytoplankton and periphyton. This process initiates the assimilation of floodplain <br />nutrients and potential productivity into aquatic food webs. Invertebrates play an influential role <br />in continuing the assimilation of potential productivity into aquatic food webs. Aquatic <br />invertebrates feed on periphyton, phytoplankton, and detritus, incorporating nutrients and <br />biomass derived from the floodplain into the riverine food web. Grazer, shredder, and filter <br />feeding invertebrates are in turn eaten by predatory invertebrates and fish, thus assimilating <br />floodplain productivity into the top levels of the food web. <br />Zooplankton and aquatic invertebrate larvae are an important food resource to juvenile <br />fish. Therefore, the high density and diversity of invertebrate assemblages within floodplain <br />habitats represents an augmented food resource to juvenile endangered fish. Because of <br />enhanced productivity of food organisms, floodplain wetlands facilitate rapid fish growth and <br />survival (Osmundson 1986) and enhance fish survivorship within the river system. Native <br />young-of-year (YOY) fish that leave the river and use floodplain wetlands should grow faster <br />i than if they had remained in the river. Larger native (YOY) fish should be less susceptible to <br />predation by most nonnative fish in the main channel. Thus, it is hypothesized that floodplain <br />restoration will provide juvenile endangered fish access to high quality nursery habitat with <br />' enhanced food resources and favorable physical conditions which should enhance survivorship. <br />These potential benefits could be offset, however, if large numbers of nonnative fish species also <br />use these habitats. <br />Floodplain habitats in the middle Green River are believed to be important to razorback <br />sucker and other native fishes (Wydoski and Wick 1998, Modde 1997). The chronology of <br />spawning and larval emergence (Tyus and Karp 1990, Modde et al. 1996, Muth 1998), as well <br />as a linkage between juvenile razorback sucker and floodplains (Modde 1996) has led some to <br />suggest that floodplain habitats are important to early life stages of razorback sucker. In <br />' response to an increasing body of data linking floodplain habitat to razorback sucker, the <br />Recovery Implementation Program (RIP) initiated a floodplain enhancement program which <br />includes floodplain acquisition (David Soker, U.S. Fish and Wildlife Service, personal <br />communication) and construction (e.g., levee removal) to increase access of floodplains to m <br />endangered fishes. The goal of this program is to enhance floodplain habitats for the benefit of <br />endangered fishes. <br />1 However, several issues have arisen that address the biological costs and benefits of : <br />floodplain enhancement. Lentsch et al. (1996) presented a levee removal strategy that outlined <br />the need to monitor natural as well as enhanced floodplain habitats and emphasized yt1le need M <br />monitor the impact of floodplain enhancement on the status of nonnative fishes in tti@'Gree& <br />2 <br />1