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Technical Report Series U.S. Fish and Wildlife Service The Fish and Wildlife Service publishes five technical report series. Manuscripts are accepted from Service employees or contractors, students and faculty associated with cooperative fish and wildlife research units, and other persons whose work is sponsored by the Service. Manuscripts are received with the understanding that they are unpublished. Most manuscripts receive anonymous peer review. The final decision to publish lies with the editor. Editorial Staff MANAGING EDrrOR Paul A. Opler ASSISTANT SECTION LEADER Paul A. Vohs WILDLIFE EDITOR Elizabeth D. Rockwell FIsIIERIEs EDrrOR James R. Zuboy PUBLICATIONS MANAGEMENT Thomas J. Cortese TEcEmcAL EDrmRS Deborah K. Harris, Senior Editor John S. Ramsey VISUAL INFORMATION SPECIALIST Constance M. Lemos EDrmRIAL ASSISTANTS Rhonda G. Brown Amy D. Trujillo Series Descriptions Biological Report ISSN 0895-1926 Technical papers about applied research of limited scope. Subjects include new information arising from comprehensive studies, surveys and inventories, effects of land use on fish and wildlife, diseases of fish and wildlife, and developments in technology. Proceedings of technical conferences and symposia may be published in this series. Fish and Wildlife Leaflet ISSN 0899-451X Summaries of technical information for readers of non- technical or semitechnical material. Subjects include topics of current interest, results of inventories and surveys, management techniques, and descriptions of imported fish and wildlife and their diseases. Fish and Wildlife Research ISSN 1040-2411 Papers on experimental research, theoretical presentations, and interpretive literature reviews. North American Fauna ISSN 0078-1304 Monographs of long-term or basic research on faunal and floral life histories, distributions, population dynamics, and taxonomy and on community eulogy. Resource Publication ISSN 0163-4801 Semitechnical and nonexperimental technical topics including surveys; data, status, and historical reports; handbooks; checklists; manuals; annotated bibliographies; and workshop papers. Copies of this publication may be obtained from the Publications Unit, U.S. Fish and Wildlife Service, 1849 C Street, N.W., Mail Stop 130, Webb Building, Washington, DC 20240, or may be purchased from the National Technical Information Service (NTIS), 5285 Port Royal Road, Springfield, VA 22161 (call toll free 1-800-553-6847).