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TIME OF ISOLATION OF CYPRINODON DIABOLIS IN DEVILS HOLE: <br />GEOLOGIC EVIDENCE <br />by Isaac J. Winograd and Barney J. Szabo <br />ABSTRACT <br />The modern water table in Devils Hole is about 16 m below <br />land surface. It is commonly believed that Cvprinodon diabolis <br />became isolated in this cavern at the close of the Pleistocene <br />Epoch, 12,000 to 15,000 years ago, as a direct result of <br />increasing aridity accompanied by a cessation of spring discharge <br />from Devils Hole. Uranium-series dating of speleothems from <br />Brown's Room in Devils Hole indicates, however, that the water <br />table stood only about 9 m above the modern level between 44,000 ± <br />1,000 and 27,000 ± 1,000 years ago, and declined thereafter. <br />Thus, if we assume that C. diabolis became isolated due to a <br />decline of water table then this species has been isolated a <br />minimum of 44,000 years. <br />What might be the maximum time of isolation of C. diabolis? <br />The distribution of vein calcite, tufa, and other evidence <br />indicative of paleo-ground water discharge, indicates that during <br />the early-to-middle Pleistocene (1.6 million to 700,000 years ago) <br />the water table in the central Amargosa Desert, Nevada was tens of <br />meters higher than the modern water table, and that ground-water <br />discharge occurred as far as 14 km upstream from modern discharge <br />areas. The apparent water-table decline in this part of the Great <br />Basin since the early-to-middle Pleistocene probably reflects a <br />combination of: a) tectonic uplift of vein calcite and tufa, <br />unaccompanied by a change in water-table altitude; b) lowering of <br />ground water base-level in response to tectonism or erosion; and; <br />c) decline in water-table altitude in response to increasing <br />aridity caused by major uplift of the Sierra Nevada and Transverse <br />Ranges during the Pleistocene. Uranium-series dating of the <br />calcitic veins permits calculation of rates of apparent water- <br />tab13 decline in the past million years; rates of 0.02 to 0.09 <br />m/10 yr are indicated. <br />A synthesis of hydrogeologic, neotectonic, and paleo- <br />climatologic information with the vein-calcite data permits the <br />inference that the water table in the central Amargosa Desert, <br />progressively lowered throughout the Pleistocene. If we assume <br />that C. diabolis was isolated due to this progressive lowering of <br />the water table at a proto-Devils Hole, then -- using the water- <br />49