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The geomorphology of the study reach was characterized using closely spaced measurements of <br />channel gradient, bed material and cross section morphology. These measurements indicate that the <br />characteristics of the Colorado River change systematically downstream, but not in the manner <br />expected from geomorphological studies of other rivers. Specifically, we observe that the grain size <br />of the substrate sediment decreases rather slowly in comparison to the average gradient. This <br />observation contrasts with the common assumption that substrate sediment size and slope are <br />highly correlated. In addition, we observe that the average bankfull depth increases downstream <br />more rapidly than the average bankfull width. Over the 300-km length of the study area, the <br />bankfull depth increases by roughly 100% (doubles) while the bankfull width increases by only <br />about 30%. This result contrasts with results from many other studies showing that the bankfull <br />width of an alluvial river typically increases downstream faster than the bankfull depth. <br />Thresholds for coarse-sediment transport were estimated on the basis of flow modeling at 10 sites, <br />and measurements of channel characteristics throughout the study area. These results indicate that <br />the framework gravels of the Colorado begin moving at flows somewhere between 35 and 55% of <br />the bankfull discharge; flows higher than this occur on average about 30 days per year. Flows <br />equal to the bankfull discharge are assumed to rework most of the bed material; the frequency of <br />these flows varies from several (5-7) days per year in upper reaches to a few (1-2) days per year in <br />lower reaches. The magnitudes of these threshold flows are shown to be similar when individual <br />reaches are grouped according to the two major tributaries in the area. For the three reaches above <br />the Gunnison River, we estimate that discharges of 211 to 278 m3/s (7500-9800 ft3/s) will initiate <br />motion, while discharges of 580 to 623 m3/s (20,500-22,000 ft3/s) will reach the bankfull level; for <br />the reaches between the Gunnison River and the Dolores River, discharges of 497 to 548 m3/s <br />(17,500-19,300 ft3/s) will initiate motion, and discharges of 979 to 1320 m3/s (34,600-46,600 ft3/s) <br />will reach the bankfull level; and for the reaches below the Dolores River, initial motion occurs at <br />discharges ranging from 561 to 659 m3/s (19,800-23,300 ft3/s), and bankfull flow occurs at <br />discharges ranging from about 1500 to 2000 m3/s (54,500-71,000 ft3/s). <br />The biological implications of our results can be summarized as follows: Mobile gravel substrates <br />exist throughout the study area, as do off-channel habitats such as backwaters. However, these <br />features vary in their relative abundance and potential suitability as habitat, depending on location <br />within the study area. Alluvial reaches above Westwater Canyon are characterized by lower flow <br />depths and somewhat coarser substrates than the reaches downstream. However, because of the <br />influence of channel slope, substrate mobility is roughly the same in all reaches, with framework <br />gravels moving at flows equal to about half the bankfull discharge and higher. These flows are an <br />important prerequisite for maintaining mobile bed features and in-channel habitats such as riffles <br />and gravel bars. Perhaps just as important are the conditions favoring primary and secondary <br />productivity. Biological processes at these lower trophic levels are affected by a variety of factors, <br />including nutrient availability, water temperature, turbidity and interstitial void space. Our results <br />indicate that suspended sediment concentrations (turbidity) are typically higher in the lower reaches <br />of the study area than they are in the upper reaches. In addition, our cross section data show that <br />the channel becomes considerably deeper downstream without getting much wider. The net effect <br />of the changes in sediment concentration and cross-section shape is a more box-like channel where <br />proportionally less and less of the bed receives sufficient sunlight to allow primary production. As <br />a result, factors that enhance primary and secondary productivity (clear, shallow water) are <br />optimized in the upper reaches whereas factors that might limit these processes (turbid, deep water) <br />become important in the lower reaches (see related reports by Lamarra, 1999 and Osmundson, 1999 <br />for complete descriptions of the aquatic community structure in the Colorado River). <br />V1 <br />