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develop physically based models for discharges that will improve existing fish habitats. The <br />specific objectives of this study were as follows: <br />1) Determine how reservoir operations have changed the annual flow hydrograph and <br />sediment-transport capacity of the upper Colorado River; <br />2) Quantify the effect of these historical changes on channel morphology;. <br />3) Measure existing channel characteristics and responses to snowmelt runoff events; and <br />4) Provide recommendations for flows that will maintain or improve existing habitats. <br />The present study focuses on a 90-km segment of the upper Colorado River from Palisade, CO to <br />Westwater, UT. This segment provides important habitat for Colorado squawfish and razorback <br />sucker and marks the upper limit of their range on the main-stem Colorado River. We also include <br />analyses of a 85-km reach of the lower Gunnison River between Delta and Grand Junction, CO. <br />In conducting this work we were fortunate to have abundant flow and sediment load data, high <br />quality aerial photographs, and several years of above-average runoff in which we could observe <br />the effects of high flows on the river. Our results not only provide key information for fisheries <br />biologists and water resource engineers, they also give added insight into questions about rates of <br />channel change, mechanisms of cross section and profile adjustment, and processes of sediment <br />transport in gravel-bed rivers. <br />RELATION BETWEEN FISH HABITATS AND GEOMORPHOLOGY <br />The present study focuses on habitats used primarily by Colorado squawfish. Of the four <br />endangered species, Colorado squawfish are perhaps the most studied, and they are certainly the <br />most abundant of the endangered species in the study area (Stanford and Ward, 1986). The <br />population of razorback sucker in the upper Colorado River is very small, and possibly no longer <br />self-sustaining (Osmundson and Kaeding, 1991). Some of the information in this report is <br />applicable to razorback sucker, but this species is considerably different from Colorado squawfish, <br />thus we restrict most of our analysis to processes and conditions that affect the latter species. <br />Ecology and Habitat Use <br />The ecology and habitat needs of Colorado squawfish have been described in detail in a number of <br />studies (reviewed by Tyus, 1991 and Stanford, 1994). It is important to note here that much of the <br />present understanding of Colorado squawfish ecology is based on studies of a relatively large <br />population in the Green-Yampa River system. Early studies of this population showed that <br />individual squawfish made long (> 100 km) seasonal migrations to spawn in specific reaches of <br />the Green and Yampa Rivers (Tyus and Karp, 1989; Tyus, 1991). Similar studies of the <br />Colorado-Gunnison River subpopulation of squawfish suggest that, while there is a tendency for <br />adults to congregate near Grand Junction during the spawning season, they migrate relatively short <br />distances (23 km, on average), and they spawn in widely separated reaches (McAda and Kaeding, <br />1991). For whatever reason, squawfish in the upper Colorado River are not as specific in their <br />selection of spawning sites as their Green-Yampa counterparts are. It does appear, however, that <br />the basic requirements for spawning are similar. Spawning occurs several weeks after the peak in <br />the snowmelt hydrograph between late June and early August when water temperatures reach 18- <br />22°C (McAda and Kaeding, 1991). In the few instances where spawning has been observed in the <br />upper Colorado River, the fish were seen congregating near dissected gravel bars formed by loose, <br />open-framework particles (D. Osmundson, personal communication), similar to what has been <br />observed on the Yampa River (Lamarra et al., 1985; Harvey et al., 1993). <br />2