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Methods <br />For these studies backwaters were trammel netted during spring runoff each year from 1991 <br />through 1994. Subadults and adults congregate in these low-velocity habitats during runoff <br />when main-channel flows increase dramatically. Fish were entrapped in backwaters by a <br />method dubbed `scare and snare'. Using a jon boat and a crew of two people, the open end of <br />each backwater was first blocked with a trammel net; the boat then entered the backwater by <br />passing over the net with the motor raised. Once inside, the boat was used to scare fish <br />toward the net by vigorously motoring back and forth beginning at the far end of the <br />backwater and working towards the mouth. Nets were pulled as soon as sufficient `scare' <br />effort was expended (5-15 min depending on backwater size). The entire range of the <br />Colorado River population upstream of the Green River confluence was systematically <br />sampled each year, with three complete sampling passes made annually through the upper <br />reach (upstream of Westwater Canyon) and two annual sampling passes made through the <br />lower reach (downstream of Westwater Canyon). With each pass, every backwater suspected <br />of harboring adult Colorado squawfish was netted. Sampling effort was augmented with <br />electrofishing in some segments where backwaters were few. Additional capture data from <br />concurrent studies were utilized to the extent possible. Data from fish recaptured during other <br />studies in 1995 were also used to supplement growth and movement data. <br />Results and Discussion <br />Growth, age and survival <br />Mean annual growth rates of 3- to-6-year-oid fish were calculated from measured lengths of <br />individuals aged by scale analysis. The highest rate was 82.0 mm/year when fish were three <br />years old. By the time fish were six years old, average growth rate declined to 32.2 mm/year. <br />Growth rates for older fish were estimated from changes in total length (TL) of marked and <br />later recaptured individuals. These annual growth increments were averaged for fish within 50 <br />mm size groups. For fisli 500-549 mm TL, annual growth increments averaged 22.2 mm/yr. <br />For fish > 550 mm TL, growth increments averaged 8.7 mm/year, but varied (although not <br />significantly) from 15.0 (700-799 mm-long fish) to 3.9 mm/year (800-899 mm-long fish). <br />Thus, rate of growth slows markedly after- about age 10, although there appears to be an <br />increase in rate when fish are between 650 and 800 mm TL; beyond 800 mm TL, growth <br />apparently slows again. For fish up to 10 years old, results here were similar to those <br />reported in previous studies. However, measured increments from capture-recapture data <br />indicated that previous scale-based methods over estimated real growth rates in older fish (> <br />10 years old). <br />Based on these new growth curves, ages offish of various length were calculated. <br />Simulations indicated broad ranges of ages for fish of given lengths, especially fish > 550 mm <br />TL: it takes 10-22 (X = 15) years for 95% of the fish to reach 600 mm in length; 16-30 (X = <br />25) years to reach 700 corn; 20-40 (X = 32)years to reach 800 mm. <br />ii