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Because pools are highly preferred during winter, weighted area also goes up. However, because <br />we do not know if backwaters, eddies and pools will still be preferred at flows less than 1,654 cfs <br />during winter, we cannot recommend a winter flow of 557 cfs based on weighted area of pools, <br />backwaters and eddies. Thus we recommend 1,630 cfs as the best discharge level for maximizing <br />the area of preferred habitats during winter. <br />Mean winter flows (November-March) during the historic period (1902-1942) range from 1,322 <br />(January) to 1,789 cfs (November) and averaged 1,531 cfs for all months (see Osmundson and <br />Kaeding 1991). Recent (1954-1989), mean, winter flows range from 1,765 (January) to 2,161 cfs <br />(November) and average 1,920 cfs. Thus, our recommended winter flow of 1,630 cfs would be <br />100 cfs higher than average historic winter flows and 290 cfs less than recent winter averages. <br />Habitat Depth <br />Minimum Flows for Providing Suitable Depth <br />Habitat quality needs to be considered along with maximizing habitat area. Two measurable <br />variables that affect quality are habitat depth and habitat diversity. Depth, considered in this <br />section, is an important component of cover. As discussed earlier, depth and turbidity are probably <br />the primary sources of cover for fishes in this portion of the Colorado River, though during winter, <br />ice provides a third cover component (Osmundson and Kaeding 1989). <br />Backwaters, eddies and pools are the preferred habitat types of adult squawfish during summer and <br />winter. During winter, pools replace eddies as the most preferred type. Depth at fish locations <br />within these habitat types varied somewhat between summer and winter (Fig. 20; Table 2). <br />Though depths at fish locations in eddies did not differ significantly between seasons, depths in <br />backwaters and pools selected by the fish did differ between summer and winter. Fish used deeper <br />backwater sites in summer than in winter and used deeper pools in winter than in summer. <br />Our objective was to use the stage/depth information obtained from the 12 transects to see if depth <br />of preferred habitats was suitable or not during the flow levels recommended from the mapping <br />output. We felt that for each habitat type, the average depth used by the fish, or rather the range of <br />variance about the mean (plus or minus one standard error), provided a reasonable estimate of <br />optimum or at least suitable depth. Preferred depth, a better indicator of optimum depth, could not <br />be determined because measuring preference requires a comparison between use and availability. <br />We had too little depth data to determine availability. Using the depth-use data, backwaters, for <br />example, would have a range of suitable depths during summer of 3.4-4.5 ft; during winter, 2.6-3.1 <br />ft (see Table 2). Presumably, water more shallow than that generally used by the fish provides <br />insufficient cover. Water deeper than average should provide additional cover and therefore is not <br />considered detrimental. Low use of deep water likely reflects its low availability. <br />Ideally, using the above example, we would like to know at what flow level the average and <br />maximum depth of most backwaters in the 15-mile reach is at least 3.4 ft during summer and 2.6 ft <br />during winter. This was the rationale behind monitoring stage at our transect sites. Stage, along <br />with the bed cross-section data, allowed us to measure maximum and average depth at various <br />flows (Table 3). However, there were not enough transects to allow us to make more general <br />conclusions about relationships between stage and depth of habitats in general. For example, to <br />37