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capacity for the reach can be expected. Aggressive, introduced carp and channel catfish may <br />become concentrated in low-velocity eddies and pools; this might explain displacement of <br />squawfish into runs. <br />In the 15-mi1e reach, squawfish have two options in dealing with low summer flow conditions. <br />One method is to `hole up' in portions of slow runs that remain relatively deep due to springtime <br />scouring of the bed and the presence of a downstream control structure such as a submerged <br />cobble bar. At such times the fish are concentrated in these sites and are not free to utilize the <br />resources of the entire river. The strategy is to wait out the low-water period until higher flows <br />return. The second strategy is for the fish to travel downstream and wait out the low-flow period in <br />reaches below the confluence with the Gunnison River where more water is available. <br />Though our sample is small, we have some evidence to indicate that adult squawfish may indeed <br />leave the 15-mile reach when water becomes low. During 1986 we maintained contact with three <br />squawfish from May through December. All three spent the year within the reach (one briefly left <br />during the spawning season and returned in July). Mean monthly flows during August, September <br />and October in 1986 were 2,040, 1,868 and 2,223 cfs, respectively. During 1987, mean flows <br />were somewhat lower: 1,214 cfs in August, 812 cfs in September and 733 cfs in October. We <br />maintained contact with 5 squawfish during 1987. Three remained in the reach year-round (one <br />left and returned in July), one left and returned by October 9 prior to the end of the irrigation <br />season, and one left and returned on November 9 shortly after the irrigation season ended and more <br />water was returned to the reach. Mean flows were lowest in 1988: 588 cfs in August, 542 cfs in <br />September and 178 in October. Contact was maintained with two squawfish; both left the reach <br />after the spawning period was over and did not return that year. One left in late September or early <br />October; the other left in late July or early August. Such displacements from the home range are <br />no doubt stressful for fish and strongly suggest a seasonal reduction in carrying capacity for the <br />reach - this in a river where quality adult habitat is already much reduced in miles. <br />During our 1986-1988 radiotelemetry study, squawfish experienced a range of summer flow levels. <br />The months of August, September and October of 1986 and August of 1987 had moderate-flows <br />comparable to historic (1902-1942) summer flow levels (see Osmundson and Kaeding 1991). <br />September and October of 1987 and 1988 and August of 1988 had flows slightly lower than the <br />mean flows of recent (1954-1989) summer months. All summer flows today are approximately <br />1,600 cfs lower than they would be under virgin conditions because of water diverted immediately <br />upstream by local irrigation companies. These withdrawals predate the historic period. Thus, <br />without alterations, a summer day with a flow of 500 cfs, for example, would be more than 2,100 <br />cfs under normal conditions. <br />Tyus (1992) warned against determining instream flow needs of fish based on fish habitat use data <br />collected under altered, suboptimal and perhaps unsuitable habitat conditions. Resulting recom- <br />mendations will likely provide a minimum amount of water needed for fish survival rather than a <br />more favorable amount needed to promote population recovery. Data from the 1986-1988 study <br />indicated that during low-water conditions some squawfish left the 15-mile reach and those that <br />stayed displayed a distinct change in habitat selection. We interpret such behavioral changes as <br />reflective of suboptimal conditions. Our view is that squawfish habitat use data collected from the <br />15-mile reach during abnormally low-water conditions may demonstrate the ability of the species to <br />modify its habitat use patterns to enable it to temporarily cope with adverse conditions, but it is not <br />32