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Movements by G. cypha within the LCR.---Our results contrast with <br />those of Kaeding and Zimmerman (1983), who found no consistent <br />relationship between catch rate and river reach within the LCR <br />(where 'river reaches' were 5 km increments, beginning at RKM 2 <br />and ending at Blue Springs (RKM 21; Fig. 1--2b)). In our <br />analyses, river reaches were more extensive, and only encompassed <br />those RKM within which G. cypha was active (i.e., 0--14.9). <br />The confluence has often been considered a staging area for <br />G. cypha (R. R. Miller, GCNP report, 1975, unpubl.; R. D. <br />Suttkus, G. H. Clemmer, C. Jones, and C. R. Shoop, GCNP report, <br />1976, unpubl.; C. 0. Minckley, unpubl. field notes, 1977). <br />Extent of G. cypha's movement within the LCR was not clarified <br />until September 1977, when three large individuals (278--295 mm <br />TL) were captured 12.8 RKM above the confluence (C. 0. Minckley, <br />unpubl. field notes, 1977). From these data, and from AZGF <br />monitoring efforts in spring 1987--1990 (C. 0. Minckley, <br />unpubl.), it was believed that G. cypha actively moved into the <br />LCR in spring (i.e., April/May) to reproduce, then quickly <br />returned to the mainstem. Greater numbers of G. cypha at the <br />confluence during spring of 1992 support an hypothesis of staging <br />prior to upstream movement. Downstream (i.e „ postreproductive) <br />movement also clearly occurred, but spanned a long period and was <br />diffuse. Movements between LCR reaches during a given sampling <br />period were negligible, suggesting temporal closure during <br />periods of sampling. There was no evidence of explosive or <br />extensive reproductive movements. <br />Our results indicate population stasis within reaches, <br />particularly summer through winter, suggesting that G. cypha is <br />more of a resident component of the LCR than previously imagined. <br />These observations support similar data collected by Karp and <br />Tyus (1990) in the Yampa River. There, G. cypha remained in or <br />near specific eddies for extended periods, and even returned to <br />the same eddy during the spawning season in different years. It <br />could not be ascertained whether individual chub deposited eggs <br />in the eddies or simply used them for staging, resting, or <br />feeding. <br />Habitat use.-- rata on habitat use by G. cypha are primarily <br />anecdotal and --,servational. Adults characterize whitewater <br />reaches, where they occupy deep, swirling eddies along canyon <br />walls or concentrate in zones of turbulence near boulders and <br />submerged rocks (Minckley 1991:150). Similarly, Kaeding et al. <br />(1990) noted that commonality among G. cypha habitats is not <br />great depth, but dynamic flow vectors that result from water <br />moving rapidly among large, angular boulders and shoreline rock <br />outcrops. Within other areas of the Colorado River, G. cypha <br />often associates with large-scale riprap material from riverside <br />railroad and highway construction (Kaeding et al., 1990). <br />Karp and Tyus (1990) argued that eddy habitat was crucial to <br />10