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also Minckley, 1983). They were also eliminated from the Green <br />River above the mouth of the Yampa River when Flaming Gorge Dam <br />became operational in 1962 (Vanicek et al., 1970; Fig. 1--2a). <br />Glen Canyon Dam.---The operation of Glen Canyon Dam precipitated <br />major changes in the Marble/Grand Canyon ecosystem of the <br />Colorado River (Marzolf, 1991:33). Some occurred immediately upon <br />closure of the dam in 1963 (e.g., decreased water temperatures; <br />reduced sediment loads; diminished salinity; alteration of flow <br />regimes). Others developed over a much longer time frame (e.g., <br />geomorphic adjustment of channel; secondary succession of <br />terrestrial vegetation; modification of aquatic species- <br />composition) (Committee, 1991). All have severely impacted the <br />natural ecosystem; some are irreversible. <br />Indigenous fishes inhabiting Glen, Marble and Grand canyons <br />were impacted following closure of Glen Canyon Dam (Holden and <br />Stalnaker, 1975; Suttkus and Clemmer, 1977; Minckley, 1991). Many <br />(including G. cypha: Holden and Stalnaker, 1975; Anonymous, 1980) <br />persisted in Lake Powell, but were unable to reproduce (Holden, <br />1973:4). Downstream from the dam, the fish community shifted from <br />predominantly warm-water native and introduced fishes to one <br />dominated by either cold-water fishes [i.e., rainbow trout <br />(Oncorhynchus mykiss) and brown trout (Salmo trutta)] or those <br />with broad temperature tolerances. Within GCNP, five of eight <br />indigenous fishes still persist in low to moderate numbers. These <br />are usually restricted to warmer habitats such as tributaries and <br />backwaters. While terrestrial species in GCNP adapted to the <br />post-dam Colorado River ecosystem (Carothers and Brown 1991:147; <br />Johnson, 1991), indigenous fishes generally did not (Kaeding and <br />Zimmerman, 1983:592). <br />The Little Colorado River as habitat.---Temperataure and flow <br />conditions in the LCR are similar to those of the pre-dam <br />Colorado mainstem, and thus suit habitat requirements of <br />indigenous fishes shaped over evolutionary time. Kaeding and <br />Zimmerman (1983) argued that G. cypha persisted within the <br />Canyon, whereas other endemics were eliminated, because a portion <br />of its population spawned within the LCR. They also argued that, <br />given post-dam temperature disparities between LCR and mainstem, <br />significant reproductive success for G. cypha must occur within <br />t`-?e LCR. Thus, selection should be strong for development of a <br />spawning migration (Kaeding and Zimmerman, 1983). Critical though <br />these observations are to the ecology and conservation of G. <br />cypha, they have yet to be substantiated. While data presented <br />herein do not address movements of G. cypha from the mainstem <br />into the LCR, they do suggest that staging occurs at the <br />confluence. Our data do demonstrate that adult G. cypha actively <br />move up the LCR in spring (primarily to reproduce), and often <br />remain within the LCR for long periods, possibly the entire year. <br />These observations are based both on monthly population estimates <br />by reach, and on seasonal recaptures of tagged G. cypha. Before <br />8