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Reproduction <br />Smallmouth bass spawn in spring, usually mid-April to July (Watson 1955; <br />Henderson and Foster 1957; Harlan and Speaker 1969; Turner and MacCrimmon <br />1970; Smitherman and Ramsey 1972; Neves 1975; Pflieger 1975), depending on <br />geographical location and water temperature. Cleary (1956) observed a 45 day <br />post-nesting period for smallmouth spawning in Iowa streams. <br />Smallmouth bass spawn on rocky lake shoals, river shallows, or backwaters <br />or move into creeks or tributaries to- spawn (Harrison 1954; Webster 1954; <br />Cleary 1956; Sanderson 1958; Harlan and Speaker 1969; Coble 1975; Clancey <br />1980). Smallmouth bass spawned in warm sloughs or backwater areas bordering <br />the Columbia River (Montgomery et al. 1980). The species requires a clean <br />stone, rock, or gravel substrate for spawning (Robbins and MacCrimmon 1974). <br />Studies show that the habitat condition during spawning is the most important <br />factor for year class strength in smallmouth bass (Clancey 1980). <br />Nest building and spawning occur when the water temperature is <br />12.8-21.0° C (Turner and MacCrimmon 1970; Scott and Crossman 1973; Shuter <br />et al. 1980), but most activity occurs at or above 15° C (Henderson and Foster <br />1957; Latta 1963; Harlan and Speaker 1969; Coble 1975; Shuter et al. 1980). <br />Specific Habitat Requirements <br />4 <br />Optimum riverine habitat is characterized by cool, clear, midorder streams <br />> 10.5 m wide (Carlander 1977) with abundant shade and cover (Larimore et al. ' <br />1952) and deep pools, moderate current, and a gravel or rubble substrate <br />(Sanderson 1958; Robbins and MacCrimmon 1974; Coble 1975; Pflieger 1975). <br />Streams with gradients of 0.75-4.70 m/km, that provide alternating pools and <br />riffles, support the largest stream populations of smallmouth bass (Trautman <br />1942; Hallam 1959; Robbins and MacCrimmon 1974; Coble 1975; Funk and Pflieger <br />1975). Standing crop is generally largest in pools deeper than 1.2 m (Robbins <br />and MacCrimmon 1974; Paragamian 1979; Clancey 1980). <br />Growth of smallmouth bass is faster in lakes and reservoirs than in <br />rivers (Robbins and MacCrimmon 1974). Optimum lacustrine habitat is char- <br />acterized by large, clear lakes and reservoirs with an average depth greater <br />than 9 m with rocky shoals (Turner and MacCrimmon 1970; Coble 1975; Miller <br />1975; Pflieger 1975). Large concentrations of smallmouth bass occurred over a <br />broken rock and boulder substrate with a large amount of interstitial space in <br />the Tongue River Reservoir in Montana (Penkal and Gregory 1980). No bass were <br />found over a rounded rock substrate or sand in the Snake River (Munther 1970). <br />Although smallmouth are able to live in small ponds (Bennett 1965; Buck and <br />Thoits 1970; Coble 1975), populations are more successful in larger lakes and <br />reservoirs (Jenkins 1975). Smallmouth are found almost exclusively in the <br />epilimnion during summer stratification in northeastern Wisconsin and Ontario <br />(Hile and Juday 1941; Turner and MacCrimmon 1970), yet frequent depths up to <br />12 m in all seasons in northern New York (Webster 1954). <br />Smallmouth bass exhibit strong, cover-seeking behavior and prefer protec- <br />tion from light in all life stages (Coble 1975; Miller 1975). Deep, dark <br />2